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big squirreltail, big squirreltail grass

diverseglume wildrye, unequal-glumed wildrye

Habit Plants cespitose, not rhizomatous. Plants cespitose, not rhizomatous, sometimes moderately glacous.
Culms

15-65 cm, erect to ascending, usually puberulent;

nodes 4-6, mostly concealed, glabrous.

70-160 cm, erect;

nodes 4-9, mostly exposed, glabrous.

Leaves

evenly distributed;

sheaths glabrous or white-villous;

auricles usually present, 0.5-1.5 mm;

ligules to 1 mm, truncate, entire or lacerate;

blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous.

evenly distributed;

sheaths glabrous, often purplish;

auricles 1-2 mm, purplish or brownish black;

ligules usually 1-2 mm;

blades 5-17 mm wide, lax, adaxial surfaces usually pilose, at least on the veins, occasionally scabrous.

Spikes

5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes;

internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets.

8-28 cm long, 3-5 cm wide, nodding to pendent, with 2 spikelets per node, rarely with 1 or 3 at a few nodes;

internodes 4-6(9) mm long, 0.2-0.3 mm thick at the thinnest sections, margins and summits often pubescent.

Spikelets

10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

10-16 mm, appressed, with 2-4(5) florets, lowest florets functional;

disarticulation above the glumes, beneath each floret.

Glumes

subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity;

fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate;

paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate;

anthers 1-2 mm.

usually differing in length by at least (3)4 mm, occasionally obsolete, (1)2-15(20) mm long including the undifferentiated awns, indurate at the base, (0.1)0.2-0.4(0.6) mm wide, setaceous, 0-1-veined, tapering from the base, scabrous or hispidulous at least towards the apices, margins firm, awns often outcurving;

lemmas 7-12 mm, usually silvery-hirsute to sericeous, occasionally hirtellous or strigose, at least near the margins, backs sometimes scabrous, awns 18-35 mm, moderately to strongly outcurving at maturity;

paleas 7-10 mm, obtuse, occasionally emarginate;

anthers 2-4 mm.

Anthesis

from late May to June.

from early June to late July.

2n

= 28.

= 28.

Elymus multisetus

Elymus diversiglumis

Distribution
from FNA
AZ; CA; CO; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; IA; IL; MI; MN; MO; ND; OK; SD; WI; WY; MB; ON; SK
[BONAP county map]
Discussion

Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric.

Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus diversiglumis grows in moist to dry, often base-rich and alluvial soils, in open woods, woodland margins, and thickets in the northern Great Plains, from Saskatchewan and Manitoba to Wyoming, Wisconsin, and Iowa.

Elymus diversiglumis is a variable species that, like E. svensonii (p. 314) and E. churchii (p. 314), may have originated from hybrids between E. canadensis var. canadensis (p. 305) and E. hystrix (see next), although part of its range extends further west than the current distribution of the latter species. Elymus diversiglumis usually reaches anthesis 2-4 weeks earlier than sympatric populations of E. canadensis. Church (1954, 1958, 1967a) found that most artificial canadensis-hystrix hybrids, as well as some plants of E. diversiglumis itself, are sterile. Those that were not sterile could occasionally form fertile backcrosses with E. canadensis and, to a lesser extent, with E. hystrix. Introgressant populations involving all three species are known. Artificial crosses with other species have not been successful.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 318. FNA vol. 24, p. 316.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Name authority (J.G. Sm.) Burtt Davy Scribn. & C.R. Ball.
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