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big squirreltail, big squirreltail grass

wildrye

Habit Plants cespitose, not rhizomatous. Plants loosely cespitose, without conspicuous rhizomes.
Culms

15-65 cm, erect to ascending, usually puberulent;

nodes 4-6, mostly concealed, glabrous.

30-130 cm tall, 1-5 mm thick, erect or weakly decumbent;

nodes 3-4, glabrous, glaucous.

Leaves

evenly distributed;

sheaths glabrous or white-villous;

auricles usually present, 0.5-1.5 mm;

ligules to 1 mm, truncate, entire or lacerate;

blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous.

basal and cauline;

sheaths glaucous, glabrous or with hairs, lower sheaths sometimes hairy, upper sheaths glabrous, margins sometimes ciliate;

auricles 1.5-2.5 mm;

ligules about 0.3 mm;

blades 10-25 cm long, 3-10 mm wide, glabrous or pilose.

Spikes

5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes;

internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets.

10-22 cm long, 1.5-2.5 cm wide including the awns, 0.8-1 cm wide excluding the awns, inclined to nodding, with 1 spikelet at all or most nodes;

rachises scabrous on the edges, glabrous below the spikelets;

internodes 10-25 mm.

Spikelets

10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

5-22 mm long, about 5 mm wide, appressed, with 4-12 florets;

disarticulation above the glumes, beneath the florets.

Glumes

subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity;

fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate;

paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate;

anthers 1-2 mm.

narrowly elliptic to lance-oblong, apices acute to acuminate;

lower glumes 5-11 mm;

upper glumes 7-13 mm;

lemmas 7-12 mm, mostly glabrous, glabrate, or sparsely hairy, margins with coarse stiff hairs, hairs to 1 mm, apices abruptly narrowed, awned, awns 10-20 mm, scabrous, strongly outcurved to recurved;

paleas 2/3 - 4/5 the length of the lemmas, keels winged distally, distinctly outwardly curved below the apices, apices 0.5-0.6 mm wide, truncate to rounded;

anthers about 2 mm.

Anthesis

from late May to June.

Genome

StY.

2n

= 28.

= 28.

Elymus multisetus

Elymus ciliaris

Distribution
from FNA
AZ; CA; CO; ID; NV; OR; UT; WA; WY
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Discussion

Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric.

Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus ciliaris is native to northern China and Japan. It was collected from ballast dumps in Portland, Oregon, in 1899 and 1902; it is not established in the Flora region. A.S. Hitchcock identified both specimens on the sheet (US 1017954) as Agropyron caninum (L.) P. Beauv. [= Elymus caninus, p. 322], from which E. ciliaris differs in its short, rounded paleas and relatively short glumes with distinctly outwardly curving keels. The other specimen on that sheet is E. tsuskushiensis (p. 336).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 318. FNA vol. 24, p. 336.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Name authority (J.G. Sm.) Burtt Davy (Trin.) Tzvelev
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