Elymus multisetus |
Elymus caninus |
|
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big squirreltail, big squirreltail grass |
bearded wheatgrass |
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Habit | Plants cespitose, not rhizomatous. | Plants cespitose, not strongly rhizomatous. |
Culms | 15-65 cm, erect to ascending, usually puberulent; nodes 4-6, mostly concealed, glabrous. |
30-130 cm, erect or geniculate, usually hairy on or below the nodes. |
Leaves | evenly distributed; sheaths glabrous or white-villous; auricles usually present, 0.5-1.5 mm; ligules to 1 mm, truncate, entire or lacerate; blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous. |
evenly distributed; sheaths glabrous; auricles to 1.5 mm; ligules 0.2-1.5 mm; blades 10-30 cm long, 4-10 mm wide, flat, both surfaces scabrous, adaxial surfaces sometimes with hairs over the veins, hairs to 0.5 mm, veins not prominent, widely spaced. |
Spikes | 5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes; internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets. |
5-20 cm long, 0.5-1.5 cm wide including the awns, 5-8 mm wide excluding the awns, erect or arching, with 1 spikelet per node; internodes 4.5-7 mm, edges scabrous or ciliate, both surfaces hairy below the spikelets. |
Spikelets | 10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node; disarticulation initially at the rachis nodes, subsequently beneath each floret. |
10-15(20) mm long, 2-5(7) mm wide, appressed to slightly divergent, with 2-6 florets; rachillas scabridulous or pubescent, often more densely pubescent distally; disarticulation above the glumes, beneath each floret. |
Glumes | subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity; fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate; paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate; anthers 1-2 mm. |
equal to unequal, 0.6-1 mm wide, lanceolate to narrowly ovate, usually green, flat or weakly keeled, keels eccentric, adaxial surfaces hairy, hairs often inconspicuous, hyaline margins sometimes widest distally, narrowing abruptly to the acute to acuminate apices; lower glumes 8-11 mm, 3-veined, usually awned, awns to 3 mm; upper glumes 10-13 mm, 3-5-veined, sometimes awn-tipped, awns to 0.3 mm; lemmas 9-13 mm, glabrous, smooth to somewhat scabridulous distally, rounded on the back proximally, awned, awns 7-20 mm, straight or flexuous; paleas subequal to the lemmas, keels finely and densely ciliate over most of their length, straight or slightly outwardly curved, tapering to the apices, apices about 0.2 mm wide; anthers 2-3 mm. |
Anthesis | from late May to June. |
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Haplomes | StH. |
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2n | = 28. |
= 28. |
Elymus multisetus |
Elymus caninus |
|
Distribution |
AZ; CA; CO; ID; NV; OR; UT; WA; WY
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OR; WA |
Discussion | Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric. Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus caninus is native to Eurasia; it is not known to be established in the Flora region. A.S. Hitchcock (1935, 1951) reported that it had been collected on ballast dumps in Portland, Oregon, but the specimens concerned belong to E. ciliaris (p. 336) and E. tsukushiensis (p. 336). Elymus caninus differs from E. ciliaris and E. tsukushiensis in having flatter glumes that are longer in relation to the lemmas, and palea keels that are straight or almost straight below the apices. Recent reports of its occurrence in the region reflect C.L. Hitchcock et al.'s (1969) treatment, in which E. caninus and E. trachycaulus were treated as conspecific subspecies. Because E. caninus is the older name, it is the correct name to use at the specific rank under such a treatment. The hairs on the inside of the glumes are difficult to see. Nevertheless, this is the single most reliable morphological character for distinguishing Elymus caninus from all other species of Elymus in this treatment. Elymus caninus is most likely to be confused with awned plants of E. trachycaulus (p. 321). The two species also differ in their molecular characteristics, and in at least one chromosome interchange (Sun et al. 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 318. | FNA vol. 24, p. 322. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus |
Sibling taxa | ||
Synonyms | Agropyron caninum | |
Name authority | (J.G. Sm.) Burtt Davy | (L.) L. |
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