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big squirreltail, big squirreltail grass

Canada wild rye, Canadian wild rye, Great Plains wild-rye, élyme du Canada

Habit Plants cespitose, not rhizomatous. Plants loosely cespitose, rarely with rhizomes to 4 cm long and 1-2 mm thick, often glaucous.
Culms

15-65 cm, erect to ascending, usually puberulent;

nodes 4-6, mostly concealed, glabrous.

(40)60-150(180) cm, erect or decumbent;

nodes 4-10, mostly concealed by the leaf sheaths, glabrous.

Leaves

evenly distributed;

sheaths glabrous or white-villous;

auricles usually present, 0.5-1.5 mm;

ligules to 1 mm, truncate, entire or lacerate;

blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous.

evenly distributed;

sheaths smooth or scabridulous, glabrous or hirsute, often reddish brown;

auricles 1.5-4 mm, brown or purplish black;

ligules to 1(2) mm, truncate, ciliolate;

blades (3)4-15(20) mm wide, usually firm, often ascending and somewhat involute, usually dull green, drying to grayish, adaxial surfaces usually smooth or scabridulous and glabrous, rarely sparsely hispid to villous.

Spikes

5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes;

internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets.

6-30 cm long, 3-7 cm wide, usually nodding, sometimes pendent or almost erect, usually with 2(3) spikelets per node, occasionally to 5 at some nodes, rarely with 1 at some nodes but never throughout;

internodes (2)3-5(7) mm long, or 5-10 mm long towards the base, 0.2-0.35 mm thick at the thinnest sections, glabrous or with a few hairs below the spikelets.

Spikelets

10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

12-20 mm excluding the awns, more or less divergent, with (2)3-5(7) florets, lowest florets functional;

disarticulation usually above the glumes and beneath each floret, rarely also below the glumes.

Glumes

subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity;

fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate;

paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate;

anthers 1-2 mm.

usually equal, occasionally subequal, 11-40 mm including the awns, the basal 0-1 mm subterete and slightly indurate, glume bodies 6-13 mm long, 0.5-1.6 mm wide, linear-lanceolate to subsetaceous, entire, widening or parallel-sided above the base, 3-5-veined, glabrous to scabrous-ciliate, rarely villous on the veins, margins firm, awns (5)10-25(27) mm, straight to outcurving;

lemmas 8-15 mm, glabrous, scabrous, hispid, or uniformly villous with the hairs generally appressed, awns (10)15-40(50) mm, moderately to strongly outcurving, often contorted at the spike bases;

paleas 7-13 mm, acute, usually bidentate;

anthers 2-3.5 mm.

Anthesis

from late May to June.

May to July.

2n

= 28.

= 28, rarely 42.

Elymus multisetus

Elymus canadensis

Distribution
from FNA
AZ; CA; CO; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric.

Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus canadensis grows on dry to moist or damp, often sandy or gravelly soil on prairies, dunes, stream banks, ditches, roadsides, and disturbed ground, or, especially to the south, in thickets and open woods near streams. It is widespread in most of temperate North America, extending from the southwestern Northwest Territories to Coahuila, Mexico, being especially common in the Great Plains. Reports from California and the southeastern states appear to be based on misidentifications. E. canadensis is considered a good forage species.

Elymus canadensis is sometimes confused with E. riparius (see previous), from which it differs in having curved rather than straight awns; and with E. wiegandii (p. 305), from which it differs in its less robust habit and narrower leaves. It can hybridize with E. glabriflorus (p. 296), E. virginicus (p. 298), E. hystrix (p. 316) and allies, E. glaucus (p. 306), E. trachycaulus (p. 321), Pseudoroegneria spicata (p. 281), and other species. Subsequent introgression may have contributed to much of the diversity within the genus (Pohl 1959; Brown and Pratt 1960; Nelson and Tyrl 1978; Davies 1980; Campbell 2002). The three varieties recognized here show clear differences in their typical expression and evidence some geographic separation, but they may prove to be artificial reference points within a more or less continuous variation (Sanders et al. 1979). Nevertheless, crossing barriers sometimes exist between the varieties, and even between some sympatric strains (Church 1954, 1958, 1967a).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas usually villous or hispid; spikes nodding to almost pendent; internodes 4-7 mm long, often strongly glaucous
var. canadensis
1. Lemmas usually smooth or scabridulous, occasionally hirsute; spikes usually nodding, occasionally almost erect; internodes 3-4 mm long, not strongly glaucous.
→ 2
2. Glumes not clearly indurate or bowed out at the base, awns 10-20 mm long; lemmas smooth or scabridulous, awns usually 20-30 mm long, moderately outcurving; spikes 6-20 cm long
var. brachystachys
2. Glumes often slightly indurate and bowed out at the base, awns 15-25 mm long; lemmas occasionally hirsute, awns 30-40 mm long, often strongly outcurving; spikes 15-25(30) cm long
var. robustus
Source FNA vol. 24, p. 318. FNA vol. 24, p. 303.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Subordinate taxa
E. canadensis var. brachystachys, E. canadensis var. canadensis, E. canadensis var. robustus
Name authority (J.G. Sm.) Burtt Davy L.
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