Elymus macgregorii
Elymus wawawaiensis
early wild-rye
Snake River wheatgrass, wawawai wild rye
40-120 cm, erect or slightly decumbent;
nodes 4-8, mostly exposed, glabrous.
(15)50-130 cm, erect, mostly glabrous;
nodes usually glabrous, sometimes slightly pubescent.
evenly distributed;
sheaths usually glabrous, rarely villous;
auricles 2-3 mm, usually purplish black when fresh, sometimes light brown;
ligules shorter than 1 mm;
blades 7-15 mm wide, lax, dark glossy green under the glaucous bloom, adaxial surfaces usually glabrous, occasionally villous.
more or less evenly distributed;
basal sheaths glabrate, margins not evidently ciliate;
auricles absent or to 1.2 mm;
ligules 0.1-1.1 mm;
blades to 28 cm long, 1.7-5 mm wide, involute when dry, adaxial surfaces usually densely pubescent, rarely sparsely pubescent.
4-12 cm long, (1.7)2.2-3(4)4 cm wide, erect, exserted, with (6)9-16(20) nodes and 2 spikelets at all or most nodes, sometimes with 3 at some nodes;
internodes 4-7 mm long, about 0.3 mm thick and 2-angled at the thinnest sections, usually glabrous or scabridulous beneath the spikelets.
5-20 cm long, 2.5-3 cm wide including the awns, erect to slightly nodding, with 1 spikelet per node;
internodes 5-12 mm long, about 0.2 mm thick, about 0.3 mm wide, glabrous beneath the spikelets.
10-15 mm, strongly divergent, glaucous, maturing to pale yellowish brown, with (2)3-4 florets, lowest florets functional;
disarticulation below the glumes and each floret, the lowest floret often falling with the glumes.
10-22 mm long, about twice as long as the internodes, 2-8.5 mm wide, appressed, with 4-10 florets;
rachillas glabrous;
disarticulation above the glumes, beneath each floret.
subequal, entire, the basal 1-3 mm terete or subterete, indurate, without evident venation, moderately bowed out, glume bodies 8-16 mm long, 1-1.8 mm wide, linear-lanceolate, widening or parallel-sided above the base, (2)4-5(8)-veined, usually glabrous, occasionally hirsute, sometimes scabrous, margins firm, awns (10)15-20(25) mm, straight except the awns of the lowest spikelets occasionally contorted;
lemmas 6-12 mm, usually glabrous, sometimes scabrous, occasionally villous, awns (15)20-30 mm, straight;
paleas 6-10 mm, apices obtuse;
anthers 2-4 mm.
4-10 mm long, 0.5-1.3 mm wide, narrowly lanceolate, widest at or below midlength, glabrous, often glaucous, 1-3-veined, flat or weakly keeled, margins 0.1-0.2 mm wide, widest near midlength, apices usually acuminate, awned or unawned, awns to 6 mm;
lemmas 6-12 mm, smooth or slightly scabrous, margins often sparsely pubescent proximally, apices awned, longest awns in the spikelets 9-28 mm, strongly divergent;
paleas 7.2-10.5 mm, keels scabrous distally, tapering to the 0.2-0.3 mm wide apices;
anthers 3.5-6 mm.
usually mid-May to mid-June.
= 28.
= 28.
Elymus macgregorii
Elymus wawawaiensis
Elymus macgregorii grows in moist, deep, alluvial or residual, calcareous or other base-rich soils in woods and thickets, mostly east of the 100th Meridian in the contiguous United States. It used to be confused with E. glabriflorus (p. 296) or E. virginicus (p. 298), but it reaches anthesis about a month earlier than sympatric populations of these species. In most of its range, E. macgregorii has purplish black auricles; light brown auricles may be locally abundant, particularly in populations at the limits of its range.
Elymus macregorii hybridizes with several species, but especially E. virginicus and E. hystrix (p. 316) (Campbell 2000). Western plants often have smaller, more condensed spikes and distinctly villous leaves, suggesting a transition to E. virginicus var. jejunus (p. 300). Transitions to E. virginicus var. jejunus can also be recognized to the north, where the dates of anthesis are delayed, but even in Maine, E. macgregorii reaches anthesis about 10 days earlier than E. virginicus (Campbell and Haines 2002). Plants with villous lemmas grow at scattered locations; they have not been reported in distinct habitats, nor in large enough populations to warrant taxonomic recognition.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus wawawaiensis grows primarily in shallow, rocky soils of slopes in coulees and reaches of the Salmon, Snake, and Yakima rivers of Washington, northern Oregon, and Idaho. There are also a few records from localities at some distance from the Snake River and its tributaries. These probably reflect deliberate introductions. C.V. Piper, who worked for the U.S. Department of Agriculture in southeastern Washington from 1892-1902, frequently distributed seed to farmers in the region from populations that he considered superior; he considered E. wawawaiensis to be a superior form of what is here called Pseudo-roegneria spicata (p. 281). Another source of introduced populations is 'Secar', a cultivar of E. wawawaiensis that is recommended as a forage grass for arid areas of the northwestern United States.
Elymus wawawaiensis resembles a vigorous version of Pseudoroegneria spicata, and was long confused with that species. It differs in its more imbricate spikelets and narrower, stiff glumes. In its primary range, E. wawawaiensis is often sympatric with P. spicata, but the two tend to grow in different habitats, E. wawawaiensis growing in shallow, rocky soils and P. spicata in medium- to fine-textured loess soil. The two species also differ cytologically, E. wawawaiensis being an allo-tetraploid, and P. spicata consisting of diploids and autotetraploids.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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