Elymus macgregorii
Elymus violaceus
early wild-rye
Alaska wildrye, arctic wheatgrass, arctic wildrye, bearded wheatgrass, high wheatgrass, élyme latiglume
40-120 cm, erect or slightly decumbent;
nodes 4-8, mostly exposed, glabrous.
18-75 cm, often decumbent or geniculate;
nodes usually glabrous.
glabrous;
auricles about 0.5 mm;
ligules 0.5-1 mm, truncate;
blades 3-4 mm wide, flat, glabrous or hairy, abaxial surfaces less densely hairy and with shorter hairs than the adaxial surfaces, apices acute.
evenly distributed;
sheaths usually glabrous, rarely villous;
auricles 2-3 mm, usually purplish black when fresh, sometimes light brown;
ligules shorter than 1 mm;
blades 7-15 mm wide, lax, dark glossy green under the glaucous bloom, adaxial surfaces usually glabrous, occasionally villous.
4-12 cm long, (1.7)2.2-3(4)4 cm wide, erect, exserted, with (6)9-16(20) nodes and 2 spikelets at all or most nodes, sometimes with 3 at some nodes;
internodes 4-7 mm long, about 0.3 mm thick and 2-angled at the thinnest sections, usually glabrous or scabridulous beneath the spikelets.
5-12 cm long, 0.4-0.7 cm wide excluding the awns, erect, with 1 spikelet per node;
internodes 4-5.5 mm, edges ciliate.
10-15 mm, strongly divergent, glaucous, maturing to pale yellowish brown, with (2)3-4 florets, lowest florets functional;
disarticulation below the glumes and each floret, the lowest floret often falling with the glumes.
11-19 mm, appressed, with (3)4-5 florets;
rachillas hairy, hairs about 0.4 mm;
disarticulation above the glumes, beneath each floret.
subequal, entire, the basal 1-3 mm terete or subterete, indurate, without evident venation, moderately bowed out, glume bodies 8-16 mm long, 1-1.8 mm wide, linear-lanceolate, widening or parallel-sided above the base, (2)4-5(8)-veined, usually glabrous, occasionally hirsute, sometimes scabrous, margins firm, awns (10)15-20(25) mm, straight except the awns of the lowest spikelets occasionally contorted;
lemmas 6-12 mm, usually glabrous, sometimes scabrous, occasionally villous, awns (15)20-30 mm, straight;
paleas 6-10 mm, apices obtuse;
anthers 2-4 mm.
8-12 mm long, 1.2-2 mm wide, about 3/4 as long as to equaling the adjacent lemmas, narrowly ovate to obovate, often purplish, glabrous, sometimes scabrous, flat or equally keeled the full length, keels and other veins usually smooth, sometimes scabrous, 3(5)-veined, adaxial surfaces glabrous, margins usually unequal, the wider margin 0.3-1 mm wide, usually widest in the distal 1/3, apices acute to rounded, often awned, awns to 2 mm;
lemmas glabrous or pubescent, hairs flexible, all similar, apices usually awned, awns 0.5-3 mm, straight;
paleas subequal to the lemmas, tapering to the apices, apices about 0.4 mm wide;
anthers 0.7-1.3 mm.
usually mid-May to mid-June.
StH.
= 28.
= 28.
Elymus macgregorii
Elymus violaceus
Elymus macgregorii grows in moist, deep, alluvial or residual, calcareous or other base-rich soils in woods and thickets, mostly east of the 100th Meridian in the contiguous United States. It used to be confused with E. glabriflorus (p. 296) or E. virginicus (p. 298), but it reaches anthesis about a month earlier than sympatric populations of these species. In most of its range, E. macgregorii has purplish black auricles; light brown auricles may be locally abundant, particularly in populations at the limits of its range.
Elymus macregorii hybridizes with several species, but especially E. virginicus and E. hystrix (p. 316) (Campbell 2000). Western plants often have smaller, more condensed spikes and distinctly villous leaves, suggesting a transition to E. virginicus var. jejunus (p. 300). Transitions to E. virginicus var. jejunus can also be recognized to the north, where the dates of anthesis are delayed, but even in Maine, E. macgregorii reaches anthesis about 10 days earlier than E. virginicus (Campbell and Haines 2002). Plants with villous lemmas grow at scattered locations; they have not been reported in distinct habitats, nor in large enough populations to warrant taxonomic recognition.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus violaceus grows in arctic, subalpine, and alpine habitats, on calcareous or dolomitic rocks, from Alaska through arctic Canada to Greenland, and south in the Rocky Mountains to southern New Mexico. In western North America, it forms intermediates with E. scribneri (p. 330), E. trachycaulus (p. 321), and E. alaskanus (p. 326). It is treated here as including E. alaskanus subsp. latiglumis [= Agropyron latiglume], E. alaskanus being restricted to plants with relatively short glumes that are often found in valleys and at lower elevations than E. violaceus. Western plants of E. violaceous tend to be more glaucous, have shorter spikes and spikelets, and more obovate glumes than plants from Greenland but, until more is known about the extent and genetic basis of the variation in and among E. violaceus, E. alaskanus, and E. trachycaulus, formal taxonomic recognition seems inappropriate.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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