Elymus macgregorii
Elymus stebbinsii
early wild-rye
Parish wheatgrass, Stebbins' wheat grass, Stebbins' wildrye
40-120 cm, erect or slightly decumbent;
nodes 4-8, mostly exposed, glabrous.
60-140 cm;
nodes glabrous or retrorsely pubescent.
evenly distributed;
sheaths usually glabrous, rarely villous;
auricles 2-3 mm, usually purplish black when fresh, sometimes light brown;
ligules shorter than 1 mm;
blades 7-15 mm wide, lax, dark glossy green under the glaucous bloom, adaxial surfaces usually glabrous, occasionally villous.
evenly distributed;
sheaths glabrous or pubescent;
auricles usually present, 0.5-2 mm;
ligules 0.3-3.5 mm, truncate to acute, sometimes long-ciliate;
blades 4-6.5 mm wide, flat or the margins involute, straight.
4-12 cm long, (1.7)2.2-3(4)4 cm wide, erect, exserted, with (6)9-16(20) nodes and 2 spikelets at all or most nodes, sometimes with 3 at some nodes;
internodes 4-7 mm long, about 0.3 mm thick and 2-angled at the thinnest sections, usually glabrous or scabridulous beneath the spikelets.
15-31 cm long, 0.4-1.5 cm wide including the awns, 0.4-0.8 cm wide excluding the awns, erect, with 1 spikelet per node;
internodes 9-27 mm long, 1-1.3 mm wide, glabrous, smooth.
10-15 mm, strongly divergent, glaucous, maturing to pale yellowish brown, with (2)3-4 florets, lowest florets functional;
disarticulation below the glumes and each floret, the lowest floret often falling with the glumes.
13-29 mm long, from shorter than to almost twice as long as the internodes, 2.5-5 mm wide, appressed, with 5-7 florets;
rachillas glabrous;
disarticulation above the glumes and beneath each floret.
subequal, entire, the basal 1-3 mm terete or subterete, indurate, without evident venation, moderately bowed out, glume bodies 8-16 mm long, 1-1.8 mm wide, linear-lanceolate, widening or parallel-sided above the base, (2)4-5(8)-veined, usually glabrous, occasionally hirsute, sometimes scabrous, margins firm, awns (10)15-20(25) mm, straight except the awns of the lowest spikelets occasionally contorted;
lemmas 6-12 mm, usually glabrous, sometimes scabrous, occasionally villous, awns (15)20-30 mm, straight;
paleas 6-10 mm, apices obtuse;
anthers 2-4 mm.
subequal, 7.5-12 mm long, 1.2-1.5 mm wide, lanceolate, widest at about mid-length, flat or rounded on the back, 5-veined, veins smooth, scabrous or just the midvein scabridulous, margins widest at about midlength, apices acute, unawned;
lemmas 9-12 mm, glabrous, sometimes scabrous, acute, unawned or awned, awns to 28 mm, straight;
paleas subequal to the lemmas, tapering, apices 0.2-0.3 mm wide;
anthers (3.5)4-7 mm.
usually mid-May to mid-June.
= 28.
= 28.
Elymus macgregorii
Elymus stebbinsii
Elymus macgregorii grows in moist, deep, alluvial or residual, calcareous or other base-rich soils in woods and thickets, mostly east of the 100th Meridian in the contiguous United States. It used to be confused with E. glabriflorus (p. 296) or E. virginicus (p. 298), but it reaches anthesis about a month earlier than sympatric populations of these species. In most of its range, E. macgregorii has purplish black auricles; light brown auricles may be locally abundant, particularly in populations at the limits of its range.
Elymus macregorii hybridizes with several species, but especially E. virginicus and E. hystrix (p. 316) (Campbell 2000). Western plants often have smaller, more condensed spikes and distinctly villous leaves, suggesting a transition to E. virginicus var. jejunus (p. 300). Transitions to E. virginicus var. jejunus can also be recognized to the north, where the dates of anthesis are delayed, but even in Maine, E. macgregorii reaches anthesis about 10 days earlier than E. virginicus (Campbell and Haines 2002). Plants with villous lemmas grow at scattered locations; they have not been reported in distinct habitats, nor in large enough populations to warrant taxonomic recognition.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus stebbinsii is restricted to California, where it grows on dry slopes, chaparral, and wooded areas, at elevations below 1600 m. It differs from other Elymus species primarily in its combination of long anthers and solitary spikelets. It is often confused with E. glaucus (p. 306) and E. trachycaulus (p. 321) with solitary spikelets. It differs from both in its longer anthers, and from most representatives of E. glaucus in its acute, but unawned, glumes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Lemmas awned, awns 8-28 mm long; lower leaf sheaths rarely pubescent; spikelets 13-22 mm long | subsp. septentrionalis |
1. Lemmas unawned or with awns to 8(12) mm long; lower leaf sheaths pubescent or glabrous; spikelets 17-29 mm long | subsp. stebbinsii |
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