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early wild-rye

drooping wildrye

Habit Plants cespitose, not rhizomatous, usually glaucous. Plants cespitose, not rhizomatous.
Culms

40-120 cm, erect or slightly decumbent;

nodes 4-8, mostly exposed, glabrous.

45-135 cm, erect or geniculately ascending, glabrous.

Sheaths

glabrous or villous;

auricles to 1.5 mm;

ligules 0.5-1.5 mm, truncate;

blades 15-30 cm long, 4-12 mm wide, sometimes villous, adaxial surfaces smooth or scabrous, primary and secondary veins alternating.

Leaves

evenly distributed;

sheaths usually glabrous, rarely villous;

auricles 2-3 mm, usually purplish black when fresh, sometimes light brown;

ligules shorter than 1 mm;

blades 7-15 mm wide, lax, dark glossy green under the glaucous bloom, adaxial surfaces usually glabrous, occasionally villous.

Spikes

4-12 cm long, (1.7)2.2-3(4)4 cm wide, erect, exserted, with (6)9-16(20) nodes and 2 spikelets at all or most nodes, sometimes with 3 at some nodes;

internodes 4-7 mm long, about 0.3 mm thick and 2-angled at the thinnest sections, usually glabrous or scabridulous beneath the spikelets.

8-30 cm long, 1-2 cm wide including the awns, 0.5-1 cm wide excluding the awns, erect or nodding, usually with 1 spikelet per node, sometimes with 2 spikelets at the lower nodes;

internodes 10-20 mm long, about 0.8 mm wide, scabrous on the margins and on the surfaces, marginal prickles larger than those on the surfaces, hirtellous just below the spikelets.

Spikelets

10-15 mm, strongly divergent, glaucous, maturing to pale yellowish brown, with (2)3-4 florets, lowest florets functional;

disarticulation below the glumes and each floret, the lowest floret often falling with the glumes.

16-30 mm, loosely appressed, with 6-8 florets;

rachilla internodes about 0.8 mm, strigose, hairs to about 0.3 mm;

disarticulation above the glumes, beneath each floret.

Glumes

subequal, entire, the basal 1-3 mm terete or subterete, indurate, without evident venation, moderately bowed out, glume bodies 8-16 mm long, 1-1.8 mm wide, linear-lanceolate, widening or parallel-sided above the base, (2)4-5(8)-veined, usually glabrous, occasionally hirsute, sometimes scabrous, margins firm, awns (10)15-20(25) mm, straight except the awns of the lowest spikelets occasionally contorted;

lemmas 6-12 mm, usually glabrous, sometimes scabrous, occasionally villous, awns (15)20-30 mm, straight;

paleas 6-10 mm, apices obtuse;

anthers 2-4 mm.

subequal, 10-18 mm long, 1.1-2 mm wide, elliptic-lanceolate, green, not keeled, 5-7-veined, veins more or less equally prominent, scabrous, apices acute to acuminate;

lemmas 10-14 mm, scabrous or puberulent dorsally, awned, awns (4)12-18 mm, straight;

paleas 3/4 as long as to slightly shorter than the lemmas, keels outwardly curved below the apices, apices 0.3-0.7 mm wide, truncate;

anthers 3-6 mm.

Anthesis

usually mid-May to mid-June.

2n

= 28.

= 28.

Elymus macgregorii

Elymus semicostatus

Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MO; MS; NC; ND; NE; NH; NY; OH; OK; PA; RI; SD; TN; TX; VA; VT; WI; WV; NS; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
GA; OR
Discussion

Elymus macgregorii grows in moist, deep, alluvial or residual, calcareous or other base-rich soils in woods and thickets, mostly east of the 100th Meridian in the contiguous United States. It used to be confused with E. glabriflorus (p. 296) or E. virginicus (p. 298), but it reaches anthesis about a month earlier than sympatric populations of these species. In most of its range, E. macgregorii has purplish black auricles; light brown auricles may be locally abundant, particularly in populations at the limits of its range.

Elymus macregorii hybridizes with several species, but especially E. virginicus and E. hystrix (p. 316) (Campbell 2000). Western plants often have smaller, more condensed spikes and distinctly villous leaves, suggesting a transition to E. virginicus var. jejunus (p. 300). Transitions to E. virginicus var. jejunus can also be recognized to the north, where the dates of anthesis are delayed, but even in Maine, E. macgregorii reaches anthesis about 10 days earlier than E. virginicus (Campbell and Haines 2002). Plants with villous lemmas grow at scattered locations; they have not been reported in distinct habitats, nor in large enough populations to warrant taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus semicostatus is native to central Asia, from Afghanistan through Pakistan to northeastern India (Sikkim). Reports of its presence in the Flora region appear to be based on misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 295. FNA vol. 24, p. 338.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms Agropyron semicostatutn
Name authority R. Brooks & J.J.N. Campb. (Nees ex Steud.) Melderis
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