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early wild-rye

northern wheatgrass, tuft wheatgrass

Habit Plants cespitose, not rhizomatous, usually glaucous. Plants cespitose, sometimes appearing weakly rhizomatous.
Culms

40-120 cm, erect or slightly decumbent;

nodes 4-8, mostly exposed, glabrous.

35-100 cm, ascending to erect;

nodes sometimes pubescent.

Sheaths

glabrous;

auricles absent;

ligules 0.5-1 mm, truncate to rounded;

blades 3-10 mm wide, flat, usually glabrous, abaxial surfaces smooth or scabridulous, adaxial surfaces scabrous.

Leaves

evenly distributed;

sheaths usually glabrous, rarely villous;

auricles 2-3 mm, usually purplish black when fresh, sometimes light brown;

ligules shorter than 1 mm;

blades 7-15 mm wide, lax, dark glossy green under the glaucous bloom, adaxial surfaces usually glabrous, occasionally villous.

Spikes

4-12 cm long, (1.7)2.2-3(4)4 cm wide, erect, exserted, with (6)9-16(20) nodes and 2 spikelets at all or most nodes, sometimes with 3 at some nodes;

internodes 4-7 mm long, about 0.3 mm thick and 2-angled at the thinnest sections, usually glabrous or scabridulous beneath the spikelets.

5-20 cm long, 0.4-0.8 cm wide, erect, with 1 spikelet per node;

internodes 7-8 mm long, about 0.5 mm wide, glabrous below the spikelets.

Spikelets

10-15 mm, strongly divergent, glaucous, maturing to pale yellowish brown, with (2)3-4 florets, lowest florets functional;

disarticulation below the glumes and each floret, the lowest floret often falling with the glumes.

12-20 mm, appressed, with 4-7 florets;

rachillas hairy, hairs 0.3-0.5 mm;

disarticulation above the glumes, beneath each floret.

Glumes

subequal, entire, the basal 1-3 mm terete or subterete, indurate, without evident venation, moderately bowed out, glume bodies 8-16 mm long, 1-1.8 mm wide, linear-lanceolate, widening or parallel-sided above the base, (2)4-5(8)-veined, usually glabrous, occasionally hirsute, sometimes scabrous, margins firm, awns (10)15-20(25) mm, straight except the awns of the lowest spikelets occasionally contorted;

lemmas 6-12 mm, usually glabrous, sometimes scabrous, occasionally villous, awns (15)20-30 mm, straight;

paleas 6-10 mm, apices obtuse;

anthers 2-4 mm.

6-10 mm long, 1/3-2/3 the length of the spikelets and to about 1/2 the length of the adjacent lemmas, 0.8-1.8 mm wide, widest at about midlength, lanceolate, flat, rounded, or symmetrically keeled, usually green or green tinged with purple, 3-4-veined, veins scabridulous, scabrous, or with hairs to 0.3 mm, usually glabrous elsewhere, margins subequal, about 0.3 mm wide, widest near midlength, apices acute, unawned or awned, awns to 1 mm;

lemmas 8-12 mm, hairy throughout or glabrous distally, hairs all alike, 0.2-0.3 mm, apices unawned or awned, awns to 7 mm, straight;

paleas subequal to the lemmas, tapering to the apices, apices about 0.8 mm wide;

anthers 1-2 mm.

Anthesis

usually mid-May to mid-June.

2n

= 28.

= 28.

Elymus macgregorii

Elymus macrourus

Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MO; MS; NC; ND; NE; NH; NY; OH; OK; PA; RI; SD; TN; TX; VA; VT; WI; WV; NS; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT
[BONAP county map]
Discussion

Elymus macgregorii grows in moist, deep, alluvial or residual, calcareous or other base-rich soils in woods and thickets, mostly east of the 100th Meridian in the contiguous United States. It used to be confused with E. glabriflorus (p. 296) or E. virginicus (p. 298), but it reaches anthesis about a month earlier than sympatric populations of these species. In most of its range, E. macgregorii has purplish black auricles; light brown auricles may be locally abundant, particularly in populations at the limits of its range.

Elymus macregorii hybridizes with several species, but especially E. virginicus and E. hystrix (p. 316) (Campbell 2000). Western plants often have smaller, more condensed spikes and distinctly villous leaves, suggesting a transition to E. virginicus var. jejunus (p. 300). Transitions to E. virginicus var. jejunus can also be recognized to the north, where the dates of anthesis are delayed, but even in Maine, E. macgregorii reaches anthesis about 10 days earlier than E. virginicus (Campbell and Haines 2002). Plants with villous lemmas grow at scattered locations; they have not been reported in distinct habitats, nor in large enough populations to warrant taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus macrourus grows on river banks and bars, lake shores, and hillsides in northwestern North America. Outside of North America, it grows across the Russian arctic, and extends south into the boreal forest. Plants growing on shifting river banks and bars often appear rhizomatous, as the lower internodes elongate in response to the disturbed substrate. Plants of E. macrourus differ from E. alaskanus (see next) in the shape of their glumes and their narrower glume margins, and from E. trachycaulus (p. 321) in their relatively short glumes and evidently hairy rachilla segments.

Three varieties of Elymus macrourus are recognized in Russian treatments. It is not clear to which, if any, of the Russian varieties North American plants belong. A circumboreal study is needed, using plants grown from seeds collected in the wild. Seeds available as E. macrourus through germplasm resources appear to be misidentified.

Elymus macrourus is one of the parents in both E. ×palmerensis (p. 340) and ×Elyhordeum pilosilemma (p. 284).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 295. FNA vol. 24, p. 324.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms Agropyron macrourum, Roegneria turuchanensis, Roegneria macroura, Agropyron sericeum
Name authority R. Brooks & J.J.N. Campb. (Turcz. ex Steud.)
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