Elymus lanceolatus
Elymus riparius
streambank wheatgrass, streamside wild rye, thick-spike wildrye, thickspike wheatgrass
eastern riverbank wild-rye, river wild-rye, riverbank wildrye, élyme des rivages
22-130 cm, erect;
nodes glabrous.
70-160 cm, erect, sometimes rooting at the lower nodes;
nodes 5-10, mostly concealed, glabrous.
often mostly basal, sometimes more evenly distributed;
sheaths glabrous or pubescent;
auricles usually present on the lower leaves, 0.5-1.5 mm;
ligules 0.1-0.5 mm, erose, sometimes ciliolate;
blades 1.5-6 mm wide, generally involute, abaxial surfaces usually glabrous, adaxial surfaces strigose, ribs subequal in size and spacing.
evenly distributed;
sheaths usually glabrous or scabridulous, often reddish brown;
auricles absent or to 2 mm, brown;
ligules shorter than 1 mm;
blades (5)8-15(25) mm wide, flat, lax, dull green, drying to grayish, adaxial surfaces glabrous or scabrous.
3.5-26 cm long, 0.5-1 cm wide, erect to slightly nodding, usually with 1 spikelet per node, sometimes with 2 at a few nodes;
internodes 3.5-15 mm long, 0.1-0.8 mm wide, glabrous or hairy.
7-25 cm long, 2-4 cm wide, nodding, exserted, usually with 2 spikelets per node, rarely with 3 at some nodes;
internodes 3-5(8) mm long, 0.2-0.35 thick at the thinnest sections, usually glabrous below the spikelets.
8-31 mm, 1.5-3 times longer than the internodes, appressed, with 3-11 florets;
rachillas glabrous or hairy, hairs to 1 mm;
disarticulation above the glumes, beneath each floret.
10-20 mm, strongly divergent, with 2-3(4) florets, lowest florets functional;
disarticulation above the glumes and beneath each floret.
subequal, 5-14 mm long, 1/2 - 3/4 the length of the adjacent lemmas, 0.7-1.3 mm wide, lanceolate, glabrous or hairy, smooth or scabrous, 3-5-veined, flat or weakly, often asymmetrically keeled, keels straight, margins narrow, tapering from the base or from beyond midlength, apices acute to acuminate, sometimes mucronate or shortly awned;
lemmas 7-12 mm, glabrous or hairy, hairs all alike, sometimes scabrous, acute to awn-tipped, awns to 2 mm, straight;
paleas about equal to the lemmas, keels straight below the apices, smooth or scabrous proximally, sometimes hairy, scabrous distally, intercostal region glabrous or with hairs, apices 0.2-0.3 mm wide;
anthers (2.5)3-6 mm.
equal or subequal, 14-30 mm including the sometimes undifferentiated awn, the basal 0.5-2 mm terete, indurate, straight or nearly so, veins not evident, glume bodies 9-17 mm long, (0.3)0.5-0.8(1) mm wide, linear-setiform, entire, widening or parallel-sided above the base, 2-3(4)-veined, usually hispidulous or scabrous, rarely glabrous, margins firm, awns (5)8-18 mm, straight;
lemmas 7-14 mm, usually hispidulous, sometimes scabrous, awns 15-35 mm, usually straight, those of the basal spikelets occasionally contorted;
paleas 6-9 mm, usually acute, sometimes obtuse to truncate, bidentate;
anthers 2-2.7 mm.
late June to late July.
= 28.
= 28.
Elymus lanceolatus
Elymus riparius
Elymus lanceolatus grows in sand and clay soils and dry to mesic habitats. It is found primarily in the western half of the Flora region, between the coastal mountains and 95° W longitude, with the exception of E. lanceolatus subsp. psammophilus, which extends around the Great Lakes. Three subspecies are recognized, primarily on the basis of their lemma and palea pubescence.
Elymus lanceolatus is primarily outcrossing, and hybridizes with several species of Triticeae. Elymus albicans (p. 334) is thought to be derived from hybridization with the awned phase of Pseudoroegneria spicata (p. 281). Judging from specimens of controlled hybrids, hybridization with E. trachycaulus (p. 321) and unawned plants of P. spicata probably occur, but would be almost impossible to detect without careful observation in the field. Experimental hybrids are partially fertile, and capable of backcrossing to either parent (Dewey 1965, 1967, 1968, 1975, 1976).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus riparius grows in moist, usually alluvial and often sandy soils in woods and thickets, usually along larger streams and occasionally along upland ditches. It is widespread in most of temperate east-central North America. It is rare in southern Ontario and Quebec, and the eastern Great Plains. It is virtually absent from the southeastern coastal plain.
Elymus riparius is relatively uniform and distinct. It is sometimes confused with E. canadensis (see next), but that species has curving awns. It hybridizes occasionally with several other taxa, especially E. virginicus var. virginicus (p. 300) and E. hystrix (p. 316), but the hybrids produce only late, depauperate spikes or none at all (e.g., Church 1958).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Lemmas densely hairy, hairs flexible, some 1 mm long or longer | subsp. psammophilus |
1. Lemmas glabrous or with stiff hairs shorter than 1 mm. | → 2 |
2. Lemmas with hairs, not scabrous | subsp. lanceolatus |
2. Lemmas smooth, sometimes scabrous distally, mosdy glabrous, sometimes the lemma margins hairy proximally | subsp. riparius |
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