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southeastern wild-rye

Canada wild rye, Canadian wild rye, Great Plains wild-rye, élyme du Canada

Habit Plants cespitose, not rhizomatous, often glaucous. Plants loosely cespitose, rarely with rhizomes to 4 cm long and 1-2 mm thick, often glaucous.
Culms

60-140 cm, erect;

nodes 6-9, mostly concealed, glabrous.

(40)60-150(180) cm, erect or decumbent;

nodes 4-10, mostly concealed by the leaf sheaths, glabrous.

Leaves

evenly distributed;

sheaths glabrous or pubescent, often reddish brown;

auricles absent or to 2 mm, usually purplish brown;

ligules shorter than 1 mm;

blades 7-15 mm wide, lax or somewhat involute and ascending, usually dull green, sometimes with a glaucous bloom, adaxial surfaces glabrous or densely short-villous.

evenly distributed;

sheaths smooth or scabridulous, glabrous or hirsute, often reddish brown;

auricles 1.5-4 mm, brown or purplish black;

ligules to 1(2) mm, truncate, ciliolate;

blades (3)4-15(20) mm wide, usually firm, often ascending and somewhat involute, usually dull green, drying to grayish, adaxial surfaces usually smooth or scabridulous and glabrous, rarely sparsely hispid to villous.

Spikes

6-20 cm long, (2) 2.5-4(5.5) cm wide, erect, exserted, with (10)18-30(36) nodes, usually with 2(3) spikelets per node, occasionally with up to 5 at some nodes;

internodes 3-5 mm long, 0.3-0.8 mm thick and usually 4-angled at the thinnest sections, glabrous or pubescent beneath the spikelets.

6-30 cm long, 3-7 cm wide, usually nodding, sometimes pendent or almost erect, usually with 2(3) spikelets per node, occasionally to 5 at some nodes, rarely with 1 at some nodes but never throughout;

internodes (2)3-5(7) mm long, or 5-10 mm long towards the base, 0.2-0.35 mm thick at the thinnest sections, glabrous or with a few hairs below the spikelets.

Spikelets

10-20 mm, strongly divergent, often reddish brown at maturity, with (2)3-5(6) florets, lowest florets functional;

disarticulation below the glumes and each floret, or the lowest floret often falling with the glumes.

12-20 mm excluding the awns, more or less divergent, with (2)3-5(7) florets, lowest florets functional;

disarticulation usually above the glumes and beneath each floret, rarely also below the glumes.

Glumes

equal or subequal, entire, the basal 1-3 mm terete, indurate, moderately bowed out, without evident venation, glume bodies 7-18 mm long, (0.7)0.9-1.7 mm wide, linear-lanceolate, widening above the base, (3)4-5(7)-veined, smooth or scabrous, sometimes hirsute, margins firm, awns (10)15-25(30) mm, straight except the awns of the lowest spikelets frequently contorted;

lemmas 6-13 mm, smooth, scabrous, or hirsute, awns (15)25-35(40) mm, straight except the awns of the lowest spikelets occasionally contorted;

paleas 6-12 mm, obtuse;

anthers 2-4 mm.

usually equal, occasionally subequal, 11-40 mm including the awns, the basal 0-1 mm subterete and slightly indurate, glume bodies 6-13 mm long, 0.5-1.6 mm wide, linear-lanceolate to subsetaceous, entire, widening or parallel-sided above the base, 3-5-veined, glabrous to scabrous-ciliate, rarely villous on the veins, margins firm, awns (5)10-25(27) mm, straight to outcurving;

lemmas 8-15 mm, glabrous, scabrous, hispid, or uniformly villous with the hairs generally appressed, awns (10)15-40(50) mm, moderately to strongly outcurving, often contorted at the spike bases;

paleas 7-13 mm, acute, usually bidentate;

anthers 2-3.5 mm.

Anthesis

usually mid-June to late July.

May to July.

2n

= 28.

= 28, rarely 42.

Elymus glabriflorus

Elymus canadensis

Distribution
from FNA
AL; AR; CT; DC; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MO; MS; NC; NH; NJ; OH; OK; SC; TN; TX; VA; VT; WV
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from FNA
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
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Discussion

Elymus glabriflorus grows on moist, damp, or dry soil in open woods, thickets, and tall grasslands, sometimes spreading into old fields and roadsides. It is found in most of the southeastern United States, extending north to Iowa, Illinois, Indiana, West Virginia, and along the Atlantic coast to Maine; it is rare north of Maryland. Anthesis is usually 2-4 weeks later than in E. virginicus (see next) and other sympatric taxa, even in Texas, where it occurs up to a month earlier than the dates given (Davies 1980).

Elymus glabriflorus varies greatly in its pubescence, but without clear taxonomic relevance. Plants that combine pubescent spikelets and, usually, pubescent leaves with somewhat shorter spikes (6-12 cm versus 9-20 cm) and lemmas (6-10 mm versus 7-13 mm) are typical on relatively dry, infertile soils, especially in hilly interior regions, and are less frequent on the southeastern coastal plain. They have been named E. glabriflorus var. australis (Scribn. & C.R. Ball) J.J.N. Campb. In contrast, glabrous to scabrous plants that are often more robust usually grow on relatively moist or damp soils of bottomlands and upland depressions.

Elymus glabriflorus is most closely related to E. macgregorii (see previous) and E. virginicus, forming occasional hybrids with both (Campbell 2000). It is sometimes confused with E. villosus (p. 302), from which it differs in having erect spikes, and glumes that are bowed out and disarticulate at maturity. It has also been confused with E. canadensis, especially E. canadensis var. robustus (p. 305), which may be derived from introgressants between the two species (Davies 1980). Hybrids with E. hystrix (p. 316) are also known, with apparent introgression at some range margins. Artificial crosses with other species failed in several cases (Church 1967a, 1967b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus canadensis grows on dry to moist or damp, often sandy or gravelly soil on prairies, dunes, stream banks, ditches, roadsides, and disturbed ground, or, especially to the south, in thickets and open woods near streams. It is widespread in most of temperate North America, extending from the southwestern Northwest Territories to Coahuila, Mexico, being especially common in the Great Plains. Reports from California and the southeastern states appear to be based on misidentifications. E. canadensis is considered a good forage species.

Elymus canadensis is sometimes confused with E. riparius (see previous), from which it differs in having curved rather than straight awns; and with E. wiegandii (p. 305), from which it differs in its less robust habit and narrower leaves. It can hybridize with E. glabriflorus (p. 296), E. virginicus (p. 298), E. hystrix (p. 316) and allies, E. glaucus (p. 306), E. trachycaulus (p. 321), Pseudoroegneria spicata (p. 281), and other species. Subsequent introgression may have contributed to much of the diversity within the genus (Pohl 1959; Brown and Pratt 1960; Nelson and Tyrl 1978; Davies 1980; Campbell 2002). The three varieties recognized here show clear differences in their typical expression and evidence some geographic separation, but they may prove to be artificial reference points within a more or less continuous variation (Sanders et al. 1979). Nevertheless, crossing barriers sometimes exist between the varieties, and even between some sympatric strains (Church 1954, 1958, 1967a).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas usually villous or hispid; spikes nodding to almost pendent; internodes 4-7 mm long, often strongly glaucous
var. canadensis
1. Lemmas usually smooth or scabridulous, occasionally hirsute; spikes usually nodding, occasionally almost erect; internodes 3-4 mm long, not strongly glaucous.
→ 2
2. Glumes not clearly indurate or bowed out at the base, awns 10-20 mm long; lemmas smooth or scabridulous, awns usually 20-30 mm long, moderately outcurving; spikes 6-20 cm long
var. brachystachys
2. Glumes often slightly indurate and bowed out at the base, awns 15-25 mm long; lemmas occasionally hirsute, awns 30-40 mm long, often strongly outcurving; spikes 15-25(30) cm long
var. robustus
Source FNA vol. 24, p. 296. FNA vol. 24, p. 303.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Subordinate taxa
E. canadensis var. brachystachys, E. canadensis var. canadensis, E. canadensis var. robustus
Synonyms E. virginicus var. glabriflorus, E. virginicus var. australis, E. virginicus forma australis, E. glabriflorus var. australis, E. australis
Name authority (Vasey ex L.H. Dewey) Scribn. & C.R. Ball L.
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