Elymus curvatus |
Elymus stebbinsii |
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awnless wild-rye, awnless wlldrye, beardless wild rye |
Parish wheatgrass, Stebbins' wheat grass, Stebbins' wildrye |
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Habit | Plants cespitose, not rhizomatous, often glaucous. | Plants cespitose or shortly rhizomatous. | ||||
Culms | 60-110 cm, stiffly erect, or the base sometimes geniculate; nodes 6-9, concealed or exposed, glabrous. |
60-140 cm; nodes glabrous or retrorsely pubescent. |
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Leaves | evenly distributed; sheaths glabrous, often reddish brown; auricles to 1 mm, sometimes absent; ligules shorter than 1 mm, ciliolate; blades 5-15 mm wide, the lower blades usually lax, shorter, narrower, and senescing earlier, the upper blades usually ascending and somewhat involute, adaxial surfaces smooth or scabridulous, occasionally scabrous. |
evenly distributed; sheaths glabrous or pubescent; auricles usually present, 0.5-2 mm; ligules 0.3-3.5 mm, truncate to acute, sometimes long-ciliate; blades 4-6.5 mm wide, flat or the margins involute, straight. |
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Spikes | 9-15 cm long, (0.5)0.7-1.3 cm wide, erect, exserted or the bases slightly sheathed, with 2 spikelets per node; internodes 2.5-4.5 mm long, about 0.25-5 mm thick at the thinnest sections, smooth or scabrous beneath the spikelets. |
15-31 cm long, 0.4-1.5 cm wide including the awns, 0.4-0.8 cm wide excluding the awns, erect, with 1 spikelet per node; internodes 9-27 mm long, 1-1.3 mm wide, glabrous, smooth. |
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Spikelets | 10-15 mm, appressed, often reddish brown at maturity, with (2)3-4(5) florets, lowest florets functional; disarticulation below the glumes and beneath the florets, or the lowest floret falling with the glumes. |
13-29 mm long, from shorter than to almost twice as long as the internodes, 2.5-5 mm wide, appressed, with 5-7 florets; rachillas glabrous; disarticulation above the glumes and beneath each floret. |
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Glumes | equal or subequal, the basal 2-3 mm terete, indurate, strongly bowed out, without evident venation, glume bodies 7-15 mm long, 1.2-2.1 mm wide, linear-lanceolate, widening above the base, 3-5-veined, usually glabrous or scabrous, occasionally hispidulous, rarely hirsute on the veins, margins firm, awns 0-3(5) mm; lemmas 6-10 mm, glabrous or scabrous, rarely hirsute, awns (0.5)1-3(4) mm, rarely 5-10 mm on the lemmas of the distal spikelets, straight; paleas 6-10 mm, obtuse, often emarginate; anthers 1.5-3 mm. |
subequal, 7.5-12 mm long, 1.2-1.5 mm wide, lanceolate, widest at about mid-length, flat or rounded on the back, 5-veined, veins smooth, scabrous or just the midvein scabridulous, margins widest at about midlength, apices acute, unawned; lemmas 9-12 mm, glabrous, sometimes scabrous, acute, unawned or awned, awns to 28 mm, straight; paleas subequal to the lemmas, tapering, apices 0.2-0.3 mm wide; anthers (3.5)4-7 mm. |
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Anthesis | late June to mid-August. |
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2n | = 28, 42. |
= 28. |
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Elymus curvatus |
Elymus stebbinsii |
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Distribution |
CA
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Discussion | Elymus curvatus grows in moist or damp soils of open forests, thickets, grasslands, ditches, and disturbed ground, especially on bottomland. It is widespread from British Columbia and Washington, through the Intermountain region and northern Rockies, to the northern Great Plains. It is infrequent or rare in the midwest, the Great Lakes region, and the northeast, and is virtually unknown in the southeast. It is similar to E. virginicus (p. 298), and has sometimes been included in that species as E. virginicus var. submuticus Hook., but is more distinct than the varieties of E. virginicus treated above. Although E. virginicus and E. curvatus overlap greatly in range, E. curvatus usually has a distinct growth form, and its anthesis is 1-2 weeks later (Brooks 1974). Its spikes range from being completely exserted, especially west of the Great Plains, to largely sheathed, especially east of the Mississippi River and in more stressed environments. This geographic trend parallels that within E. virginicus, but sheathed plants of E. curvatus can usually be distinguished by their short awns. Clear transitions to E. virginicus, usually var. jejunus, are rare, but, especially from Missouri to Wisconsin, there are occasional plants with 5-10 mm awns on a few lemmas, especially at the spike tips. Rarely, plants from Missouri and Iowa to Quebec have hispid to hirsute spikelets, suggesting introgression with E. virginicus var. intermedius. There are a few records of apparent hybrids with other species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus stebbinsii is restricted to California, where it grows on dry slopes, chaparral, and wooded areas, at elevations below 1600 m. It differs from other Elymus species primarily in its combination of long anthers and solitary spikelets. It is often confused with E. glaucus (p. 306) and E. trachycaulus (p. 321) with solitary spikelets. It differs from both in its longer anthers, and from most representatives of E. glaucus in its acute, but unawned, glumes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 300. | FNA vol. 24, p. 329. | ||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. virginicus var. submuticus, E. virginicus var. jenkinsii, E. submuticus | Agropyron parishii | ||||
Name authority | Piper | Gould | ||||
Web links |