Elymus curvatus
Elymus hirsutus
awnless wild-rye, awnless wlldrye, beardless wild rye
boreal wild rye, hairy wildrye, northern ryegrass, northwestern wildrye
60-110 cm, stiffly erect, or the base sometimes geniculate;
nodes 6-9, concealed or exposed, glabrous.
40-140 cm, usually somewhat decumbent;
nodes 4-7, mostly exposed, usually glabrous, occasionally puberulent.
evenly distributed;
sheaths glabrous, often reddish brown;
auricles to 1 mm, sometimes absent;
ligules shorter than 1 mm, ciliolate;
blades 5-15 mm wide, the lower blades usually lax, shorter, narrower, and senescing earlier, the upper blades usually ascending and somewhat involute, adaxial surfaces smooth or scabridulous, occasionally scabrous.
evenly distributed;
sheaths usually glabrous and smooth, occasionally scabridulous or retrorsely hairy, sometimes purplish;
auricles to 1.5 mm, often absent;
ligules to 1 mm;
blades 4-12 mm wide, lax, usually deep green, adaxial surfaces usually pilose or villous, occasionally puberulent or scabridulous.
9-15 cm long, (0.5)0.7-1.3 cm wide, erect, exserted or the bases slightly sheathed, with 2 spikelets per node;
internodes 2.5-4.5 mm long, about 0.25-5 mm thick at the thinnest sections, smooth or scabrous beneath the spikelets.
6-20 cm long, 0.5-2 cm wide, nodding to pendent, with 2 spikelets per node, rarely with 3 at some nodes;
internodes 3-10(12) mm long, 0.2-0.7 mm thick at the thinnest sections, usually glabrous, sometimes sparsely hairy.
10-15 mm, appressed, often reddish brown at maturity, with (2)3-4(5) florets, lowest florets functional;
disarticulation below the glumes and beneath the florets, or the lowest floret falling with the glumes.
12-20 mm, appressed to divergent, sometimes purplish at higher latitudes, with 2-4(7) florets, lowest florets functional;
disarticulation above the glumes and beneath each floret.
equal or subequal, the basal 2-3 mm terete, indurate, strongly bowed out, without evident venation, glume bodies 7-15 mm long, 1.2-2.1 mm wide, linear-lanceolate, widening above the base, 3-5-veined, usually glabrous or scabrous, occasionally hispidulous, rarely hirsute on the veins, margins firm, awns 0-3(5) mm;
lemmas 6-10 mm, glabrous or scabrous, rarely hirsute, awns (0.5)1-3(4) mm, rarely 5-10 mm on the lemmas of the distal spikelets, straight;
paleas 6-10 mm, obtuse, often emarginate;
anthers 1.5-3 mm.
equal or subequal, the bases flat, occasionally indurate for 0.5 mm, veins usually evident, glume bodies (4.5)7-10(11) mm long, 0.7-1.5 mm wide, linear-lanceolate, entire, widening or parallel-sided above the base, 3-5-veined, usually scabridulous to scabrous, veins occasionally hirsute beyond midlength, margins hyaline or scarious, awns 1-10 mm, straight;
lemmas 7-14 mm, smooth or scabridulous, lateral veins hairy, margins hairy beyond midlength, marginal hairs 0.5-1 mm, longer than those elsewhere, awns (2)8-30 mm, flexuous to moderately outcurving;
paleas 6-13 mm, with hairs of varying lengths on the keels and apices, acute, bidentate;
anthers 2-3.5 mm.
late June to mid-August.
from May to July.
= 28, 42.
= 28.
Elymus curvatus
Elymus hirsutus
Elymus curvatus grows in moist or damp soils of open forests, thickets, grasslands, ditches, and disturbed ground, especially on bottomland. It is widespread from British Columbia and Washington, through the Intermountain region and northern Rockies, to the northern Great Plains. It is infrequent or rare in the midwest, the Great Lakes region, and the northeast, and is virtually unknown in the southeast. It is similar to E. virginicus (p. 298), and has sometimes been included in that species as E. virginicus var. submuticus Hook., but is more distinct than the varieties of E. virginicus treated above. Although E. virginicus and E. curvatus overlap greatly in range, E. curvatus usually has a distinct growth form, and its anthesis is 1-2 weeks later (Brooks 1974). Its spikes range from being completely exserted, especially west of the Great Plains, to largely sheathed, especially east of the Mississippi River and in more stressed environments. This geographic trend parallels that within E. virginicus, but sheathed plants of E. curvatus can usually be distinguished by their short awns. Clear transitions to E. virginicus, usually var. jejunus, are rare, but, especially from Missouri to Wisconsin, there are occasional plants with 5-10 mm awns on a few lemmas, especially at the spike tips. Rarely, plants from Missouri and Iowa to Quebec have hispid to hirsute spikelets, suggesting introgression with E. virginicus var. intermedius. There are a few records of apparent hybrids with other species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus hirsutus grows in moist to damp or dry soils in woods, thickets, and grasslands. Its range extends along the coastal mountains from the Aleutian Islands to northern Oregon, and inland to eastern British Columbia. Plants in the southern part of the range tend to have villous leaves and more erect spikes with shorter, straighter awns.
Elymus hirsutus is similar to E. glaucus (see previous), but its more pendent spikes, lemma pubescence pattern, and shorter glumes enable most specimens to be readily identified. Intermediates do exist; it is not known whether they reflect introgression or extremes of variation. Elymus hirsutus occasionally hybridizes with Leymus mollis (p. 356) and Hordeum brachyantherum (p. 243).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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