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Canada wild rye, Canadian wild rye, Great Plains wild-rye, élyme du Canada

Sierra wheatgrass, Sierra wildrye

Habit Plants loosely cespitose, rarely with rhizomes to 4 cm long and 1-2 mm thick, often glaucous. Plants cespitose, not rhizomatous.
Culms

(40)60-150(180) cm, erect or decumbent;

nodes 4-10, mostly concealed by the leaf sheaths, glabrous.

20-50 cm, prostrate or decumbent and geniculate;

nodes 1-2, exposed, glabrous.

Leaves

evenly distributed;

sheaths smooth or scabridulous, glabrous or hirsute, often reddish brown;

auricles 1.5-4 mm, brown or purplish black;

ligules to 1(2) mm, truncate, ciliolate;

blades (3)4-15(20) mm wide, usually firm, often ascending and somewhat involute, usually dull green, drying to grayish, adaxial surfaces usually smooth or scabridulous and glabrous, rarely sparsely hispid to villous.

basally concentrated;

sheaths glabrous;

auricles usually present, to 1 mm on the lower leaves;

ligules 0.2-0.5 mm, erose;

blades 1-5 mm wide, flat, abaxial surfaces smooth, glabrous, adaxial surfaces prominently ridged over the veins, with scattered hairs, hairs to 0.2 mm, veins closely spaced.

Spikes

6-30 cm long, 3-7 cm wide, usually nodding, sometimes pendent or almost erect, usually with 2(3) spikelets per node, occasionally to 5 at some nodes, rarely with 1 at some nodes but never throughout;

internodes (2)3-5(7) mm long, or 5-10 mm long towards the base, 0.2-0.35 mm thick at the thinnest sections, glabrous or with a few hairs below the spikelets.

5-15 cm long, 1.5-2.5 cm wide including the awns, 0.7-1.2 cm wide excluding the awns, flexuous, erect to nodding distally, with 1 spikelet at most nodes, occasionally some of the lower nodes with 2 spikelets;

internodes 5-15 mm long, 0.2-0.5 mm wide, both surfaces glabrous, edges ciliate, not scabrous.

Spikelets

12-20 mm excluding the awns, more or less divergent, with (2)3-5(7) florets, lowest florets functional;

disarticulation usually above the glumes and beneath each floret, rarely also below the glumes.

15-20 mm, ascending to divergent, with 3-7 florets;

rachillas glabrous;

disarticulation above the glumes, beneath each floret.

Glumes

usually equal, occasionally subequal, 11-40 mm including the awns, the basal 0-1 mm subterete and slightly indurate, glume bodies 6-13 mm long, 0.5-1.6 mm wide, linear-lanceolate to subsetaceous, entire, widening or parallel-sided above the base, 3-5-veined, glabrous to scabrous-ciliate, rarely villous on the veins, margins firm, awns (5)10-25(27) mm, straight to outcurving;

lemmas 8-15 mm, glabrous, scabrous, hispid, or uniformly villous with the hairs generally appressed, awns (10)15-40(50) mm, moderately to strongly outcurving, often contorted at the spike bases;

paleas 7-13 mm, acute, usually bidentate;

anthers 2-3.5 mm.

subequal, 6-9 mm long, 0.7-1 mm wide, lanceolate, glabrous, the bases evidently veined, apices entire, tapering into a 3-10 mm awn;

lemmas 12-16 mm, glabrous, sometimes scabridulous, apices bidentate, awned, awns 15-30 mm, arcuately diverging to strongly recurved;

paleas subequal to the lemmas, apices about 0.4 mm wide;

anthers 2-3.5 mm.

Anthesis

May to July.

2n

= 28, rarely 42.

= 28.

Elymus canadensis

Elymus sierrae

Distribution
from FNA
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from USDA
[BONAP county map]
Discussion

Elymus canadensis grows on dry to moist or damp, often sandy or gravelly soil on prairies, dunes, stream banks, ditches, roadsides, and disturbed ground, or, especially to the south, in thickets and open woods near streams. It is widespread in most of temperate North America, extending from the southwestern Northwest Territories to Coahuila, Mexico, being especially common in the Great Plains. Reports from California and the southeastern states appear to be based on misidentifications. E. canadensis is considered a good forage species.

Elymus canadensis is sometimes confused with E. riparius (see previous), from which it differs in having curved rather than straight awns; and with E. wiegandii (p. 305), from which it differs in its less robust habit and narrower leaves. It can hybridize with E. glabriflorus (p. 296), E. virginicus (p. 298), E. hystrix (p. 316) and allies, E. glaucus (p. 306), E. trachycaulus (p. 321), Pseudoroegneria spicata (p. 281), and other species. Subsequent introgression may have contributed to much of the diversity within the genus (Pohl 1959; Brown and Pratt 1960; Nelson and Tyrl 1978; Davies 1980; Campbell 2002). The three varieties recognized here show clear differences in their typical expression and evidence some geographic separation, but they may prove to be artificial reference points within a more or less continuous variation (Sanders et al. 1979). Nevertheless, crossing barriers sometimes exist between the varieties, and even between some sympatric strains (Church 1954, 1958, 1967a).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus sierrae is best known from rocky slopes and ridgetops in the Sierra Nevada, at 2100-3400 m, and is also found in Washington and Oregon. It resembles E. scribneri (see previous), differing in its non-disarticulating rachises, longer rachis internodes, and longer anthers. Hybrids with E. elymoides (p. 318) have glumes with awns 15+ mm long, and some spikelets with narrower glume bases and shorter anthers. Specimens with wide-margined glumes suggest hybridization with E. violaceus (p. 324).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas usually villous or hispid; spikes nodding to almost pendent; internodes 4-7 mm long, often strongly glaucous
var. canadensis
1. Lemmas usually smooth or scabridulous, occasionally hirsute; spikes usually nodding, occasionally almost erect; internodes 3-4 mm long, not strongly glaucous.
→ 2
2. Glumes not clearly indurate or bowed out at the base, awns 10-20 mm long; lemmas smooth or scabridulous, awns usually 20-30 mm long, moderately outcurving; spikes 6-20 cm long
var. brachystachys
2. Glumes often slightly indurate and bowed out at the base, awns 15-25 mm long; lemmas occasionally hirsute, awns 30-40 mm long, often strongly outcurving; spikes 15-25(30) cm long
var. robustus
Source FNA vol. 24, p. 303. FNA vol. 24, p. 332.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Subordinate taxa
E. canadensis var. brachystachys, E. canadensis var. canadensis, E. canadensis var. robustus
Synonyms Agropyron pringlei
Name authority L. Gould
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