Elymus canadensis |
Elymus caninus |
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Canada wild rye, Canadian wild rye, Great Plains wild-rye, élyme du Canada |
bearded wheatgrass |
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Habit | Plants loosely cespitose, rarely with rhizomes to 4 cm long and 1-2 mm thick, often glaucous. | Plants cespitose, not strongly rhizomatous. | ||||||||
Culms | (40)60-150(180) cm, erect or decumbent; nodes 4-10, mostly concealed by the leaf sheaths, glabrous. |
30-130 cm, erect or geniculate, usually hairy on or below the nodes. |
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Leaves | evenly distributed; sheaths smooth or scabridulous, glabrous or hirsute, often reddish brown; auricles 1.5-4 mm, brown or purplish black; ligules to 1(2) mm, truncate, ciliolate; blades (3)4-15(20) mm wide, usually firm, often ascending and somewhat involute, usually dull green, drying to grayish, adaxial surfaces usually smooth or scabridulous and glabrous, rarely sparsely hispid to villous. |
evenly distributed; sheaths glabrous; auricles to 1.5 mm; ligules 0.2-1.5 mm; blades 10-30 cm long, 4-10 mm wide, flat, both surfaces scabrous, adaxial surfaces sometimes with hairs over the veins, hairs to 0.5 mm, veins not prominent, widely spaced. |
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Spikes | 6-30 cm long, 3-7 cm wide, usually nodding, sometimes pendent or almost erect, usually with 2(3) spikelets per node, occasionally to 5 at some nodes, rarely with 1 at some nodes but never throughout; internodes (2)3-5(7) mm long, or 5-10 mm long towards the base, 0.2-0.35 mm thick at the thinnest sections, glabrous or with a few hairs below the spikelets. |
5-20 cm long, 0.5-1.5 cm wide including the awns, 5-8 mm wide excluding the awns, erect or arching, with 1 spikelet per node; internodes 4.5-7 mm, edges scabrous or ciliate, both surfaces hairy below the spikelets. |
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Spikelets | 12-20 mm excluding the awns, more or less divergent, with (2)3-5(7) florets, lowest florets functional; disarticulation usually above the glumes and beneath each floret, rarely also below the glumes. |
10-15(20) mm long, 2-5(7) mm wide, appressed to slightly divergent, with 2-6 florets; rachillas scabridulous or pubescent, often more densely pubescent distally; disarticulation above the glumes, beneath each floret. |
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Glumes | usually equal, occasionally subequal, 11-40 mm including the awns, the basal 0-1 mm subterete and slightly indurate, glume bodies 6-13 mm long, 0.5-1.6 mm wide, linear-lanceolate to subsetaceous, entire, widening or parallel-sided above the base, 3-5-veined, glabrous to scabrous-ciliate, rarely villous on the veins, margins firm, awns (5)10-25(27) mm, straight to outcurving; lemmas 8-15 mm, glabrous, scabrous, hispid, or uniformly villous with the hairs generally appressed, awns (10)15-40(50) mm, moderately to strongly outcurving, often contorted at the spike bases; paleas 7-13 mm, acute, usually bidentate; anthers 2-3.5 mm. |
equal to unequal, 0.6-1 mm wide, lanceolate to narrowly ovate, usually green, flat or weakly keeled, keels eccentric, adaxial surfaces hairy, hairs often inconspicuous, hyaline margins sometimes widest distally, narrowing abruptly to the acute to acuminate apices; lower glumes 8-11 mm, 3-veined, usually awned, awns to 3 mm; upper glumes 10-13 mm, 3-5-veined, sometimes awn-tipped, awns to 0.3 mm; lemmas 9-13 mm, glabrous, smooth to somewhat scabridulous distally, rounded on the back proximally, awned, awns 7-20 mm, straight or flexuous; paleas subequal to the lemmas, keels finely and densely ciliate over most of their length, straight or slightly outwardly curved, tapering to the apices, apices about 0.2 mm wide; anthers 2-3 mm. |
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Anthesis | May to July. |
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Haplomes | StH. |
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2n | = 28, rarely 42. |
= 28. |
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Elymus canadensis |
Elymus caninus |
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Distribution |
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
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OR; WA |
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Discussion | Elymus canadensis grows on dry to moist or damp, often sandy or gravelly soil on prairies, dunes, stream banks, ditches, roadsides, and disturbed ground, or, especially to the south, in thickets and open woods near streams. It is widespread in most of temperate North America, extending from the southwestern Northwest Territories to Coahuila, Mexico, being especially common in the Great Plains. Reports from California and the southeastern states appear to be based on misidentifications. E. canadensis is considered a good forage species. Elymus canadensis is sometimes confused with E. riparius (see previous), from which it differs in having curved rather than straight awns; and with E. wiegandii (p. 305), from which it differs in its less robust habit and narrower leaves. It can hybridize with E. glabriflorus (p. 296), E. virginicus (p. 298), E. hystrix (p. 316) and allies, E. glaucus (p. 306), E. trachycaulus (p. 321), Pseudoroegneria spicata (p. 281), and other species. Subsequent introgression may have contributed to much of the diversity within the genus (Pohl 1959; Brown and Pratt 1960; Nelson and Tyrl 1978; Davies 1980; Campbell 2002). The three varieties recognized here show clear differences in their typical expression and evidence some geographic separation, but they may prove to be artificial reference points within a more or less continuous variation (Sanders et al. 1979). Nevertheless, crossing barriers sometimes exist between the varieties, and even between some sympatric strains (Church 1954, 1958, 1967a). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus caninus is native to Eurasia; it is not known to be established in the Flora region. A.S. Hitchcock (1935, 1951) reported that it had been collected on ballast dumps in Portland, Oregon, but the specimens concerned belong to E. ciliaris (p. 336) and E. tsukushiensis (p. 336). Elymus caninus differs from E. ciliaris and E. tsukushiensis in having flatter glumes that are longer in relation to the lemmas, and palea keels that are straight or almost straight below the apices. Recent reports of its occurrence in the region reflect C.L. Hitchcock et al.'s (1969) treatment, in which E. caninus and E. trachycaulus were treated as conspecific subspecies. Because E. caninus is the older name, it is the correct name to use at the specific rank under such a treatment. The hairs on the inside of the glumes are difficult to see. Nevertheless, this is the single most reliable morphological character for distinguishing Elymus caninus from all other species of Elymus in this treatment. Elymus caninus is most likely to be confused with awned plants of E. trachycaulus (p. 321). The two species also differ in their molecular characteristics, and in at least one chromosome interchange (Sun et al. 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 303. | FNA vol. 24, p. 322. | ||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | ||||||||
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Synonyms | Agropyron caninum | |||||||||
Name authority | L. | (L.) L. | ||||||||
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