Elymus canadensis |
Elymus alaskanus |
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Canada wild rye, Canadian wild rye, Great Plains wild-rye, élyme du Canada |
Alaska wild rye, alaskan wheatgrass, ellisia |
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Habit | Plants loosely cespitose, rarely with rhizomes to 4 cm long and 1-2 mm thick, often glaucous. | Plants cespitose or weakly rhizomatous. | ||||||||||||
Culms | (40)60-150(180) cm, erect or decumbent; nodes 4-10, mostly concealed by the leaf sheaths, glabrous. |
20-90 cm, sometimes decumbent at the base, ascending to erect above; nodes usually pubescent, sometimes glabrous. |
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Leaves | evenly distributed; sheaths smooth or scabridulous, glabrous or hirsute, often reddish brown; auricles 1.5-4 mm, brown or purplish black; ligules to 1(2) mm, truncate, ciliolate; blades (3)4-15(20) mm wide, usually firm, often ascending and somewhat involute, usually dull green, drying to grayish, adaxial surfaces usually smooth or scabridulous and glabrous, rarely sparsely hispid to villous. |
sometimes basally concentrated; sheaths smooth or scabrous, glabrous or pilose; auricles absent or to 0.5 mm; ligules 0.2-1 mm, erose, ciliolate; blades 3-7 mm wide, flat, both surfaces smooth, scabrous, or pubescent. |
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Spikes | 6-30 cm long, 3-7 cm wide, usually nodding, sometimes pendent or almost erect, usually with 2(3) spikelets per node, occasionally to 5 at some nodes, rarely with 1 at some nodes but never throughout; internodes (2)3-5(7) mm long, or 5-10 mm long towards the base, 0.2-0.35 mm thick at the thinnest sections, glabrous or with a few hairs below the spikelets. |
3.5-14 cm long, 0.5-0.8 cm wide, erect or nodding distally, usually with 1 spikelet per node, occasionally with 2 at the lower nodes; internodes 3-10 mm long, 0.5-0.8 mm wide, mostly glabrous and smooth, edges scabrous or ciliate. |
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Spikelets | 12-20 mm excluding the awns, more or less divergent, with (2)3-5(7) florets, lowest florets functional; disarticulation usually above the glumes and beneath each floret, rarely also below the glumes. |
9-15(20) mm, 2-5 times longer than the internodes, appressed, with 3-6 florets, rachillas hispidulous; disarticulation above the glumes, beneath each floret. |
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Glumes | usually equal, occasionally subequal, 11-40 mm including the awns, the basal 0-1 mm subterete and slightly indurate, glume bodies 6-13 mm long, 0.5-1.6 mm wide, linear-lanceolate to subsetaceous, entire, widening or parallel-sided above the base, 3-5-veined, glabrous to scabrous-ciliate, rarely villous on the veins, margins firm, awns (5)10-25(27) mm, straight to outcurving; lemmas 8-15 mm, glabrous, scabrous, hispid, or uniformly villous with the hairs generally appressed, awns (10)15-40(50) mm, moderately to strongly outcurving, often contorted at the spike bases; paleas 7-13 mm, acute, usually bidentate; anthers 2-3.5 mm. |
4-8 mm long, (1.2)1.5-2 mm wide, 1/3-2/3 as long as the adjacent lemmas, oblanceolate to obovate, flat, usually purplish, glabrous or hairy, hairs 0.3-0.5 mm, margins unequal, the widest margin 0.4-1 mm wide, both margins widest above the middle, apices unawned or awned, awns to 1 mm; lemmas 7-11 mm, glabrous or hairy, sometimes scabridulous, sometimes more densely hairy distally, hairs 0.2-0.6 mm, all alike, apices unawned or awned, awns to 7 mm, straight; paleas subequal to the lemmas, keels straight below the apices; anthers 1-2 mm. |
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Anthesis | May to July. |
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2n | = 28, rarely 42. |
= 28. |
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Elymus canadensis |
Elymus alaskanus |
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Distribution |
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
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AK; CO; ID; MI; MT; NM; NV; OR; WA; WY; AB; BC; MB; NF; NS; NT; NU; ON; QC; YT; Greenland |
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Discussion | Elymus canadensis grows on dry to moist or damp, often sandy or gravelly soil on prairies, dunes, stream banks, ditches, roadsides, and disturbed ground, or, especially to the south, in thickets and open woods near streams. It is widespread in most of temperate North America, extending from the southwestern Northwest Territories to Coahuila, Mexico, being especially common in the Great Plains. Reports from California and the southeastern states appear to be based on misidentifications. E. canadensis is considered a good forage species. Elymus canadensis is sometimes confused with E. riparius (see previous), from which it differs in having curved rather than straight awns; and with E. wiegandii (p. 305), from which it differs in its less robust habit and narrower leaves. It can hybridize with E. glabriflorus (p. 296), E. virginicus (p. 298), E. hystrix (p. 316) and allies, E. glaucus (p. 306), E. trachycaulus (p. 321), Pseudoroegneria spicata (p. 281), and other species. Subsequent introgression may have contributed to much of the diversity within the genus (Pohl 1959; Brown and Pratt 1960; Nelson and Tyrl 1978; Davies 1980; Campbell 2002). The three varieties recognized here show clear differences in their typical expression and evidence some geographic separation, but they may prove to be artificial reference points within a more or less continuous variation (Sanders et al. 1979). Nevertheless, crossing barriers sometimes exist between the varieties, and even between some sympatric strains (Church 1954, 1958, 1967a). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus alaskanus extends across the high arctic of North America to extreme eastern Russia. This treatment interprets E. alaskanus as having relatively short glumes, in accordance with its treatment by Hulten (1968). Large specimens resemble E. macrourus (see previous), but differ in the shape of their glumes and in their wider glume margins. Elymus alaskanus differs from E. trachycaulus (p. 321) in its greater cold tolerance and the distal widening of its glume margins. There is some intergradation, particularly with E. violaceus (p. 324) and E. trachycaulus, but these species have longer glumes. Moreover, in western North America, E. violaceus is restricted to rocky habitats at or above treeline, whereas E. alaskanus is often associated with valleys and flat areas. Reports of its extending to New Mexico are based on the inclusion of high-elevation forms of E. trachycaulus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 303. | FNA vol. 24, p. 326. | ||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | ||||||||||||
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Synonyms | Roegneria villosa, Roegneria borealis, E. alaskanus subsp. borealis, Agropyron alaskanum | |||||||||||||
Name authority | L. | (Scribn. & Merr.) Á.Löve | ||||||||||||
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