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two-leaf waterweed

ditchmoss, water-weed

Habit Plants perennial, of fresh waters.
Rhizomes

and stolons absent.

Leaves

sometimes in 2s, spreading, linear to narrowly elliptic, 4.7–24.8 × (0.8–)1.8–4.3 mm, flat.

cauline, whorled, 3–7 at each node, or leaves opposite at 4+ proximalmost nodes, submersed, sessile;

blade linear to linear-lanceolate, apex acute;

midvein without lacunae along side(s), blade uniform in color throughout;;

abaxial surfacely without prickles or aerenchyma;

intravaginal squamules entire.

Inflorescences

staminate spathes cylindric, 10.2–42 mm;

peduncles abscissing following anthesis; pistillate spathes linear, 9–67 mm.

solitary, sessile;

spathes not winged.

Flowers

unisexual; staminate flowers: stamens 7–9;

pedicels remaining briefly following anthesis;

inner 3 filaments connate proximally, forming column;

anthers 3–4.5 mm;

pollen in monads; pistillate flowers: styles 2.3–3 mm.

unisexual, staminate and pistillate on different plants, or rarely bisexual, usually projected to surface of water by elongate floral tube base, sessile;

petals white.

Staminate flowers

filaments distinct or 3 inner connate ½ their length;

anthers oval;

pollen in monads or tetrads.

Pistillate flowers

ovary 1-locular;

styles 3, not 2-fid.

Fruits

ovoid to lance-ellipsoid, smooth, dehiscing irregularly.

Seeds

ellipsoid, 2.8–3 mm, densely covered with long hairs.

cylindric to fusiform, glabrous to hirsute.

Erect

stems rooted in substrate, branched or unbranched, elongate.

Elodea bifoliata

Elodea

Phenology Flowering summer–fall.
Habitat Waters of rivers and reservoirs
Elevation 100–2500 m (300–8200 ft)
Distribution
from FNA
CO; ID; KS; MN; MT; ND; NM; OR; SD; UT; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Central America; South America; Europe
[BONAP county map]
Discussion

An additional name, Elodea schweinitzii (G. Planchon) Caspary, has appeared in North American literature. Although it was accepted by some (H. St. John 1965), others stated, "Elodea schweinitzii is something of a taxonomist's nightmare" (C. D. K. Cook and K. Urmi-König 1985). The few collections referable to the name are exceedingly variable, so variable in fact that the authors added, "the main character is variability itself." I am following Cook and Urmi-König in not accepting the taxon.

Species 5 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Staminate spathes 4 mm or less; styles usually 2 mm or less; leaves usually less than 1.7 mm wide.
E. nuttallii
1. Staminate spathes 6 mm or more; styles usually more than 2 mm; leaves usually more than 1.8 mm wide.
→ 2
2. Pollen in tetrads; staminate pedicels detaching before or during anthesis; anthers 3 mm or less; leaves mostly in 3s; seeds 4.5–5.7 mm or more.
E. canadensis
2. Pollen in monads; staminate pedicels remaining briefly following anthesis; anthers 3–4.5 mm; some leaves in 2s; seeds 4 mm or less.
E. bifoliata
Source FNA vol. 22. FNA vol. 22.
Parent taxa Hydrocharitaceae > Elodea Hydrocharitaceae
Sibling taxa
E. canadensis, E. nuttallii
Subordinate taxa
E. bifoliata, E. canadensis, E. nuttallii
Synonyms E. longivaginata
Name authority H. St. John: Research Studies of the State College of Washington 30(2): 23. (1962) Michaux: Flora Boreali-Americana 1: 20. (1803)
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