Eleocharis uniglumis
Eleocharis rostellata
creeping spike-rush, one-glumed spikesedge, onescale spikerush, slender spike-rush, éléocharide unigume
beak spike-rush, beak spikesedge, walking sedge, walking spikerush
terete, often with some blunt ridges when dry, (5–)10–60 cm × 0.2–1.5 mm, firm, internally spongy.
1.5–3 times as wide as thick, 20–100 cm × 0.35–2 mm, firm to hard, wiry, with to 8 subacute ribs, rarely nearly smooth;
some culms arching or decumbent and rooting at tips.
distal leaf sheaths persistent, not splitting, proximally red, distally stramineous to green, often callose, thinly papery to thickly membranous, apex often dark red-brown, obtuse to subacute, tooth absent.
distal leaf sheaths not splitting abaxially, proximally dark red to brown, apex usually reddish.
ovoid to lanceoloid, 5–10 × 2–3(–4) mm, apex acute;
proximal scale amplexicaulous, entire;
subproximal scale with flower;
floral scales often spreading in fruit, 10–20, 3–4 per mm of rachilla, brown to often red-brown, midrib regions mostly stramineous to green, broadly ovate, 3–4 × 1.8–2.5 mm, entire, apex acute to obtuse, often some carinate in distal part of spikelet.
ovoid, 5–17 × 2.5–5 mm, apex acute;
spikelets on stolons rudimentary, non-flowering, proliferous when rooting;
proximal scale empty, amplexicaulous, ovate, 2–4 mm;
subproximal scale with flower;
floral scales 20–40, 2–3 per mm of rachilla, stramineous to medium brown, midrib region paler, ovate, 3.5–6 × 2–3 mm, membranous to cartilaginous, apex entire, rounded to subacute.
perianth bristles 0–4(–5), light brown to stramineous, stout, usually unequal, rudimentary to equaling achene;
stamens 3;
anthers dark yellow to stramineous, 1.2–2 mm;
styles 2-fid.
perianth bristles brown, equaling achene or tubercle, densely spinulose;
anthers brown, 2–2.4 mm.
not persistent, dark yellow or medium or dark brown, ellipsoid, obovoid, or obpyriform, biconvex, angles obscure, 1.3–1.8 × 1–1.4 mm, apex rounded, neck absent or short, smooth at 30X, or sometimes finely rugulose at 10–20X with 20 or more horizontal ridges in vertical series.
often very variable within one plant, ovoid to obovoid or obpyriform, 1.5–2.5 × 1–1.2 mm, beak to 1 × 0.6 mm.
brown to whitish, pyramidal, much higher than wide to slightly depressed, sometimes spongy and with vertical rows of depressions, 0.4–0.8 × 0.3–0.8 mm.
when present pale to dark brown, pyramidal, to 0.5 × 0.3 mm.
Eleocharis uniglumis
Eleocharis rostellata
Plants treated as Eleocharis uniglumis fall within the large morphologic variation of Eurasian E. uniglumis. Two subspecies and 3 varieties were recognized for northern Europe (S.-O. Strandhede 1966), and five species that were recognized by I. D. Zinserling (1935) were placed in synonymy under E. uniglumis (S.-O. Strandhede 1966). Recognition of infraspecific taxa within North American E. uniglumis is premature. Plants commonly called E. halophila or E. uniglumis var. halophila, found mostly in Atlantic Coastal brackish habitats, have floral scales that are usually narrower and more densely placed on the rachilla than plants usually called E. uniglumis, which are found mostly in the interior; some plants are intermediate in expression of these characters. The achene and tubercle shape characters used by M. L. Fernald (1950) to distinguish E. uniglumis from E. halophila are not valid. In North America, E. uniglumis is difficult to separate from E. erythropoda and E. kamtschatica, in both of which the spikelets have only the proximal scale without a flower (empty) and the proximal scale completely amplexicaulous. Eleocharis uniglumis differs from E. erythropoda only in its broader floral scales, which are less densely placed on the rachilla; it differs from E. kamtschatica only in its smaller tubercles. It is also difficult to separate from some specimens of E. macrostachya in which the spikelets have proximal scales that are sometimes completely amplexicaulous; such plants differ from E. uniglumis only in the absence of a flower in the axil of the subproximal scale of some of the spikelets. I have not seen voucher specimens for the chromosome numbers of 2n = 27 and 28 reported by S.-O. Strandhede (1967) from Massachusetts and Nebraska, which are lower than the 2n = (44–)46(47–88) reported for Europe (S.-O. Strandhede 1965).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Eleocharis rostellata is highly competitive, often forming large monospecific colonies. The South American E. platypus C. B. Clarke is often treated as a synonym of E. rostellata. Eleocharis rostellata superfically closely resembles E. suksdorfiana in its culms, spikelets, and achenes, but differs in the absence of creeping rhizomes, presence of stoloniferous culms, absence of a flower in the proximal scale, and achene surface details. The collection of E. rostellata I have seen from Miami-Dade County, Florida, is from 1877. I have not seen vouchers for Archuleta County, Colorado, by H. D. Harrington (1954), or for the localities in Montana and South Carolina, which are based on the map in H. K. Svenson (1934).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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