Eleocharis uniglumis
Eleocharis ravenelii
creeping spike-rush, one-glumed spikesedge, onescale spikerush, slender spike-rush, éléocharide unigume
terete, often with some blunt ridges when dry, (5–)10–60 cm × 0.2–1.5 mm, firm, internally spongy.
terete, when dry with to 12 blunt ribs, 20–55 cm × 0.5–1.1 mm, soft to firm, internally mostly hollow with complete transverse septa 2–4 mm apart, evident only on sectioning culm.
distal leaf sheaths persistent, not splitting, proximally red, distally stramineous to green, often callose, thinly papery to thickly membranous, apex often dark red-brown, obtuse to subacute, tooth absent.
distal leaf sheaths persistent, not splitting, proximally dark red, distally stramineous or reddish, thinly papery, apex often red to brown, obtuse to subtruncate, slightly callose, tooth present, 0.5–1(–3.7) mm.
ovoid to lanceoloid, 5–10 × 2–3(–4) mm, apex acute;
proximal scale amplexicaulous, entire;
subproximal scale with flower;
floral scales often spreading in fruit, 10–20, 3–4 per mm of rachilla, brown to often red-brown, midrib regions mostly stramineous to green, broadly ovate, 3–4 × 1.8–2.5 mm, entire, apex acute to obtuse, often some carinate in distal part of spikelet.
lanceoloid, 5–13 × 2–2.5 mm, apex acute to obtuse;
proximal scale amplexicaulous, entire;
subproximal scale empty;
floral scales appressed in fruit, 10–100, 10–12 per mm of rachilla, medium brown to stramineous, midrib regions often greenish, ovate, 1–1.5 × 1 mm, entire, apex rounded to subacute, carinate in distal part of spikelet.
perianth bristles 0–4(–5), light brown to stramineous, stout, usually unequal, rudimentary to equaling achene;
stamens 3;
anthers dark yellow to stramineous, 1.2–2 mm;
styles 2-fid.
perianth bristles 5–6, pale brown, stout, equaling achene;
stamens 3;
anthers brown, 0.4–0.9 mm;
styles 3-fid or a few 2-fid in the same spikelet.
not persistent, dark yellow or medium or dark brown, ellipsoid, obovoid, or obpyriform, biconvex, angles obscure, 1.3–1.8 × 1–1.4 mm, apex rounded, neck absent or short, smooth at 30X, or sometimes finely rugulose at 10–20X with 20 or more horizontal ridges in vertical series.
falling with scales, green or medium or dark brown, obpyriform, compressed-trigonous or some biconvex in same spikelet, angles prominent, 0.6–0.8 × 0.5–0.6 mm, neck short or absent, smooth or very finely reticulate at 20–30X.
brown to whitish, pyramidal, much higher than wide to slightly depressed, sometimes spongy and with vertical rows of depressions, 0.4–0.8 × 0.3–0.8 mm.
brown, depressed-pyramidal, 0.1–0.2 × 0.2–0.3 mm.
Eleocharis uniglumis
Eleocharis ravenelii
Plants treated as Eleocharis uniglumis fall within the large morphologic variation of Eurasian E. uniglumis. Two subspecies and 3 varieties were recognized for northern Europe (S.-O. Strandhede 1966), and five species that were recognized by I. D. Zinserling (1935) were placed in synonymy under E. uniglumis (S.-O. Strandhede 1966). Recognition of infraspecific taxa within North American E. uniglumis is premature. Plants commonly called E. halophila or E. uniglumis var. halophila, found mostly in Atlantic Coastal brackish habitats, have floral scales that are usually narrower and more densely placed on the rachilla than plants usually called E. uniglumis, which are found mostly in the interior; some plants are intermediate in expression of these characters. The achene and tubercle shape characters used by M. L. Fernald (1950) to distinguish E. uniglumis from E. halophila are not valid. In North America, E. uniglumis is difficult to separate from E. erythropoda and E. kamtschatica, in both of which the spikelets have only the proximal scale without a flower (empty) and the proximal scale completely amplexicaulous. Eleocharis uniglumis differs from E. erythropoda only in its broader floral scales, which are less densely placed on the rachilla; it differs from E. kamtschatica only in its smaller tubercles. It is also difficult to separate from some specimens of E. macrostachya in which the spikelets have proximal scales that are sometimes completely amplexicaulous; such plants differ from E. uniglumis only in the absence of a flower in the axil of the subproximal scale of some of the spikelets. I have not seen voucher specimens for the chromosome numbers of 2n = 27 and 28 reported by S.-O. Strandhede (1967) from Massachusetts and Nebraska, which are lower than the 2n = (44–)46(47–88) reported for Europe (S.-O. Strandhede 1965).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Of conservation concern.
Eleocharis ravenelii is apparently very uncommon in North America. It is often mistaken for very slender-stemmed E. montana, which differs in its spikelets with floral scales 1.5 mm or more, 100–500 per spikelet and 15–40 per mm of rachilla, its mostly larger and biconvex achenes, and its culm septa usually evident without sectioning the culm. Although the holotype of E. ravenelii (NY), from Corpus Christi, Texas, lacks culm bases and leaf sheaths, its culms, spikelets, floral scales, and achenes are typical of the later E. austrotexana M. C. Johnston.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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