Eleocharis uniglumis |
Eleocharis interstincta |
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creeping spike-rush, one-glumed spikesedge, onescale spikerush, slender spike-rush, éléocharide unigume |
knotted spikerush |
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Habit | Plants perennial, mat-forming; rhizomes evident, long, 0.3–1 mm thick, soft to firm, cortex often fugaceous, longer internodes 10–25 mm, scales fugaceous, 5–6 mm, membranous, not fibrous. | Plants perennial; rhizomes 2–4 mm thick, firm, longer internodes 3–7.5 cm, scales 5 mm, tubers absent. |
Culms | terete, often with some blunt ridges when dry, (5–)10–60 cm × 0.2–1.5 mm, firm, internally spongy. |
terete, 45–100 cm × (3.2–)5–9.4 mm, soft to firm, sometimes septate-nodulose proximally, not distally, internally hollow with complete transverse septa, closer together near the spikelet, evident externally; plants never forming filiform, flaccid culms. |
Leaves | distal leaf sheaths persistent, not splitting, proximally red, distally stramineous to green, often callose, thinly papery to thickly membranous, apex often dark red-brown, obtuse to subacute, tooth absent. |
distal leaf sheaths persistent, membranous to thinly papery, apex acute to acuminate. |
Spikelets | ovoid to lanceoloid, 5–10 × 2–3(–4) mm, apex acute; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales often spreading in fruit, 10–20, 3–4 per mm of rachilla, brown to often red-brown, midrib regions mostly stramineous to green, broadly ovate, 3–4 × 1.8–2.5 mm, entire, apex acute to obtuse, often some carinate in distal part of spikelet. |
not proliferous, (20–)40–62 × 4–7 mm; rachilla joints bearing prominent wing-like remnants of floral scales; proximal scale empty, amplexicaulous, (3–)3.5–5 mm; floral scales 115–220, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, obovate to broadly oblong, (4–)4.5–5 × 2.8–4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to subacute. |
Flowers | perianth bristles 0–4(–5), light brown to stramineous, stout, usually unequal, rudimentary to equaling achene; stamens 3; anthers dark yellow to stramineous, 1.2–2 mm; styles 2-fid. |
perianth bristles 6–8, stramineous, stout, flattened, subequal, exceeding achene, to 2.9 mm, coarsely spinulose; anthers stramineous to reddish, 2.5–5 mm; styles 2-fid or 3-fid. |
Achenes | not persistent, dark yellow or medium or dark brown, ellipsoid, obovoid, or obpyriform, biconvex, angles obscure, 1.3–1.8 × 1–1.4 mm, apex rounded, neck absent or short, smooth at 30X, or sometimes finely rugulose at 10–20X with 20 or more horizontal ridges in vertical series. |
stramineous to golden-yellow or reddish brown or gray, obovoid to obpyriform, biconvex or nearly plano-convex, often with abaxial longitudinal ridge, 1.4–1.8(–2) × 1.1–1.4 mm, markedly sculptured at 10–15X, each face with 23–37 rows of transversely elongated cells, the longitudinal walls separating the cells often prominent, apex with short neck 0.7–0.8 mm wide. |
Tubercles | brown to whitish, pyramidal, much higher than wide to slightly depressed, sometimes spongy and with vertical rows of depressions, 0.4–0.8 × 0.3–0.8 mm. |
dark brown, lamelliform, slightly higher than wide, 0.7–1.1 × 0.5–0.7 mm. |
Eleocharis uniglumis |
Eleocharis interstincta |
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Phenology | Fruiting summer. | Fruiting late spring–winter. |
Habitat | Mostly coastal, brackish (to fresh?) shores, marshes | Fresh ponds, lakeshores, marshes, springs, ditches, canals |
Elevation | 0–2300 m (0–7500 ft) | 10–500 m (0–1600 ft) |
Distribution |
CO; DE; MA; ME; NC; ND; NE; NH; NJ; NM; NV; NY; RI; SD; UT; VA; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; Eurasia
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FL; OK; TX; Mexico; Central America; e South America
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Discussion | Plants treated as Eleocharis uniglumis fall within the large morphologic variation of Eurasian E. uniglumis. Two subspecies and 3 varieties were recognized for northern Europe (S.-O. Strandhede 1966), and five species that were recognized by I. D. Zinserling (1935) were placed in synonymy under E. uniglumis (S.-O. Strandhede 1966). Recognition of infraspecific taxa within North American E. uniglumis is premature. Plants commonly called E. halophila or E. uniglumis var. halophila, found mostly in Atlantic Coastal brackish habitats, have floral scales that are usually narrower and more densely placed on the rachilla than plants usually called E. uniglumis, which are found mostly in the interior; some plants are intermediate in expression of these characters. The achene and tubercle shape characters used by M. L. Fernald (1950) to distinguish E. uniglumis from E. halophila are not valid. In North America, E. uniglumis is difficult to separate from E. erythropoda and E. kamtschatica, in both of which the spikelets have only the proximal scale without a flower (empty) and the proximal scale completely amplexicaulous. Eleocharis uniglumis differs from E. erythropoda only in its broader floral scales, which are less densely placed on the rachilla; it differs from E. kamtschatica only in its smaller tubercles. It is also difficult to separate from some specimens of E. macrostachya in which the spikelets have proximal scales that are sometimes completely amplexicaulous; such plants differ from E. uniglumis only in the absence of a flower in the axil of the subproximal scale of some of the spikelets. I have not seen voucher specimens for the chromosome numbers of 2n = 27 and 28 reported by S.-O. Strandhede (1967) from Massachusetts and Nebraska, which are lower than the 2n = (44–)46(47–88) reported for Europe (S.-O. Strandhede 1965). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Contrary to statements in the literature, sectioning reveals that the culm septa are closer together near the spikelet than in the rest of the culm in both Eleocharis interstincta and E. equisetoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 76. | FNA vol. 23, p. 119. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Limnochloa |
Sibling taxa | ||
Synonyms | Scirpus uniglumis, E. halophila, E. uniglumis var. halophila | Scirpus interstinctus |
Name authority | (Link) Schultes: Mant. 2: 88. (1824) | (Vahl) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 149. (1817) |
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