Eleocharis tricostata |
Eleocharis torticulmis |
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three-angle spikerush, three-angle spikesedge |
California twisted spikerush, twist-stem spikerush |
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Habit | Plants perennial, mat-forming; rhizomes evident, fairly long, 1.5–3 mm thick, hard, cortex persistent, longer internodes 1.5–3 mm, scales disintegrating to fibers, 4–5 mm, papery. | Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick. |
Culms | terete, often with 8–12 blunt ridges when dry, 20–55 cm × 0.3–1.3 mm, soft to hard. |
erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm; culm tufts not proximally bulbous. |
Leaves | distal leaf sheaths persistent, rarely splitting, proximally red to stramineous, distally green to reddish, papery, apex often dark reddish, subtruncate to obtuse, not callose, tooth present on some or all culms of plant, to 0.5 mm. |
distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse. |
Spikelets | cylindric to ovoid, 5–20 × 2–3 mm, apex rounded to subacute; proximal scale amplexicaulous, apex entire; subproximal scale empty; floral scales appressed in fruit, 20–200+, 10 per mm of rachilla, dark brown, midrib regions often stramineous or greenish, oblong, 1.7–2.5 × 1 mm, apex entire, broadly rounded, usually recurved, mostly carinate. |
6–8 × 2–3 mm; proximal scale empty, 3–4 mm, shorter than to equaling spikelet; floral scales 8–10 per spikelet, 3.5–5 × 2 mm. |
Flowers | perianth bristles absent; stamens 3; anthers dark yellow, 1–1.5 mm; styles 3-fid. |
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Perianth | bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose. |
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Achenes | falling with scales, medium to dark brown, obovoid to obpyriform, with short neck, thickly trigonous, angles prominent (keeled), 0.7–0.9 × 0.4–0.6 mm, obscurely to clearly rugulose at 10–20X with more than 20 horizontal ridges in vertical series and/or minutely cancellate at 20–30X. |
stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm; beak 0.3–0.6 mm. |
Tubercles | brown, rudimentary, mostly apiculate, sometimes pyramidal, 0.05–0.15 × 0.15–0.2 mm. |
0.25–0.6 × 0.3–0.55 mm. |
Anthers | 1.8 –3 mm. |
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Eleocharis tricostata |
Eleocharis torticulmis |
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Phenology | Fruiting late spring–summer. | Fruiting summer (Jun–Jul). |
Habitat | Wet sandy or peaty depressions, pond margins, pine barrens, savannas, mostly coastal plains | Fens, wet meadows, vernal ponds |
Elevation | 10–200 m (0–700 ft) | 1100 m (3600 ft) |
Distribution |
AL; FL; GA; MA; MI; NC; NJ; NY; SC; VA
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CA |
Discussion | I have not seen specimens to verify reports of Eleocharis tricostata from Louisiana. In the absence of rhizomes and achenes, E. tricostata is easily mistaken for E. montevidensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 88. | FNA vol. 23, p. 115. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Zinserlingia |
Sibling taxa | ||
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York. 3: 310. (1836) | S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001) |
Web links |