Eleocharis tricostata |
Eleocharis pachycarpa |
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three-angle spikerush, three-angle spikesedge |
black sand spikerush, broad fruit spikerush |
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Habit | Plants perennial, mat-forming; rhizomes evident, fairly long, 1.5–3 mm thick, hard, cortex persistent, longer internodes 1.5–3 mm, scales disintegrating to fibers, 4–5 mm, papery. | Plants perennial, forming dense clumps, often stoloniferous. |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
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Culms | terete, often with 8–12 blunt ridges when dry, 20–55 cm × 0.3–1.3 mm, soft to hard. |
erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
Leaves | distal leaf sheaths persistent, rarely splitting, proximally red to stramineous, distally green to reddish, papery, apex often dark reddish, subtruncate to obtuse, not callose, tooth present on some or all culms of plant, to 0.5 mm. |
distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
Spikelets | cylindric to ovoid, 5–20 × 2–3 mm, apex rounded to subacute; proximal scale amplexicaulous, apex entire; subproximal scale empty; floral scales appressed in fruit, 20–200+, 10 per mm of rachilla, dark brown, midrib regions often stramineous or greenish, oblong, 1.7–2.5 × 1 mm, apex entire, broadly rounded, usually recurved, mostly carinate. |
basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
Flowers | perianth bristles absent; stamens 3; anthers dark yellow, 1–1.5 mm; styles 3-fid. |
perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | falling with scales, medium to dark brown, obovoid to obpyriform, with short neck, thickly trigonous, angles prominent (keeled), 0.7–0.9 × 0.4–0.6 mm, obscurely to clearly rugulose at 10–20X with more than 20 horizontal ridges in vertical series and/or minutely cancellate at 20–30X. |
stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
Tubercles | brown, rudimentary, mostly apiculate, sometimes pyramidal, 0.05–0.15 × 0.15–0.2 mm. |
stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
Eleocharis tricostata |
Eleocharis pachycarpa |
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Phenology | Fruiting late spring–summer. | Fruiting late spring–summer. |
Habitat | Wet sandy or peaty depressions, pond margins, pine barrens, savannas, mostly coastal plains | Fresh shores, streambeds, groundwater seeps |
Elevation | 10–200 m (0–700 ft) | 100–1400 m (300–4600 ft) |
Distribution |
AL; FL; GA; MA; MI; NC; NJ; NY; SC; VA
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CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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Discussion | I have not seen specimens to verify reports of Eleocharis tricostata from Louisiana. In the absence of rhizomes and achenes, E. tricostata is easily mistaken for E. montevidensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 88. | FNA vol. 23, p. 91. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York. 3: 310. (1836) | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) |
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