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three-angle spikerush, three-angle spikesedge

ambiguous spikesedge, éléocharide ambiquë

Habit Plants perennial, mat-forming; rhizomes evident, fairly long, 1.5–3 mm thick, hard, cortex persistent, longer internodes 1.5–3 mm, scales disintegrating to fibers, 4–5 mm, papery. Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 1–3 cm, scales persistent, 5–10 mm, membranous, not fibrous.
Culms

terete, often with 8–12 blunt ridges when dry, 20–55 cm × 0.3–1.3 mm, soft to hard.

terete, often with to 16 blunt ridges when dry, 25–80 cm × 0.5–1.5 mm, firm, internally spongy.

Leaves

distal leaf sheaths persistent, rarely splitting, proximally red to stramineous, distally green to reddish, papery, apex often dark reddish, subtruncate to obtuse, not callose, tooth present on some or all culms of plant, to 0.5 mm.

distal leaf sheaths persistent, not splitting, proximally dark red, distally red to stramineous or green, often callose, thinly papery (to membranous), apex dark red-brown, obtuse to subtruncate, tooth sometimes present on some culms, to 0.3 mm.

Spikelets

cylindric to ovoid, 5–20 × 2–3 mm, apex rounded to subacute;

proximal scale amplexicaulous, apex entire;

subproximal scale empty;

floral scales appressed in fruit, 20–200+, 10 per mm of rachilla, dark brown, midrib regions often stramineous or greenish, oblong, 1.7–2.5 × 1 mm, apex entire, broadly rounded, usually recurved, mostly carinate.

ovoid to lanceoloid, 5–23 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 10–60, 3–4 per mm of rachilla, medium brown, rarely red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, in distal part lanceolate, 2.5–3.5 × 1.7 mm, entire, apex acute to rarely obtuse, often carinate in distal part of spikelet.

Flowers

perianth bristles absent;

stamens 3;

anthers dark yellow, 1–1.5 mm;

styles 3-fid.

perianth bristles 2–4(–5), brown, slender to stout, usually very unequal, rudimentary to equaling achene;

stamens 3;

anthers brown, 1.5–2 mm;

styles 2-fid, sometimes some 3-fid.

Achenes

falling with scales, medium to dark brown, obovoid to obpyriform, with short neck, thickly trigonous, angles prominent (keeled), 0.7–0.9 × 0.4–0.6 mm, obscurely to clearly rugulose at 10–20X with more than 20 horizontal ridges in vertical series and/or minutely cancellate at 20–30X.

not persistent, dark yellow or stramineous, obovoid to obpyriform, all biconvex or some (very rarely all) compressedtrigonous, angles obscure, 1–1.5 × 0.9–1.25 mm, apex rounded, neck absent (to very short), finely rugulose at 10–20X, with 20 or more horizontal ridges in vertical series and/or finely cancellate at 10–30X.

Tubercles

brown, rudimentary, mostly apiculate, sometimes pyramidal, 0.05–0.15 × 0.15–0.2 mm.

brown or proximally whitish, pyramidal, depressed, rarely some as high as wide, 0.15–0.3(–0.5) × (0.3–)0.4–0.6 mm.

2n

= 44, 45, 46.

Eleocharis tricostata

Eleocharis ambigens

Phenology Fruiting late spring–summer. Fruiting spring–summer.
Habitat Wet sandy or peaty depressions, pond margins, pine barrens, savannas, mostly coastal plains Coastal (rarely inland) fresh to brackish pond shores, marshes, disturbed places
Elevation 10–200 m (0–700 ft) 0–10 m (0–0 ft)
Distribution
from FNA
AL; FL; GA; MA; MI; NC; NJ; NY; SC; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; FL; GA; LA; MA; MS; NC; NJ; RI; SC; TX; VA; QC
Discussion

I have not seen specimens to verify reports of Eleocharis tricostata from Louisiana. In the absence of rhizomes and achenes, E. tricostata is easily mistaken for E. montevidensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis ambigens is usually treated as a synonym of E. fallax because some specimens from Massachusetts to Georgia have some 3-fid styles and some obscurely trigonous achenes. A specimen of E. ambigens from Louisiana has trigonous achenes. Eleocharis ambigens is very similar to and perhaps better included in E. uniglumis (S.-O. Strandhede 1967). Intermediates with E. macrostachya occur in Louisiana and Texas. The depressed tubercles are diagnostic of E. ambigens. Eleocharis ambigens is sometimes confused with E. montevidensis, which is readily distinguished by its floral scales with broadly rounded apices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 88. FNA vol. 23, p. 78.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York. 3: 310. (1836) Fernald: Rhodora 37: 394. (1935)
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