Eleocharis rostellata |
Eleocharis tuberculosa |
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beak spike-rush, beak spikesedge, walking sedge, walking spikerush |
cone-cup spikerush, long-tubercled spikerush, long-tubercled spikesedge |
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Habit | Plants densely tufted, mat-forming by means of rooting culm tips. | Plants perennial, forming dense clumps, not stoloniferous. |
Rhizomes | present or not, caudexlike, ascending, 1 mm thick; internodes and scales hidden by crowded culms, not evident. |
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Culms | 1.5–3 times as wide as thick, 20–100 cm × 0.35–2 mm, firm to hard, wiry, with to 8 subacute ribs, rarely nearly smooth; some culms arching or decumbent and rooting at tips. |
elliptic or sometimes circular, with several rounded ridges and sulcate when dry, 15–75 cm × 0.3(–1.5) mm, firm, minutely granular at 10X. |
Leaves | distal leaf sheaths not splitting abaxially, proximally dark red to brown, apex usually reddish. |
distal leaf sheaths persistent; stramineous to green, often minutely red-spotted, papery to membranous, apex subacute to narrowly acute, often with tooth-like callus. |
Spikelets | ovoid, 5–17 × 2.5–5 mm, apex acute; spikelets on stolons rudimentary, non-flowering, proliferous when rooting; proximal scale empty, amplexicaulous, ovate, 2–4 mm; subproximal scale with flower; floral scales 20–40, 2–3 per mm of rachilla, stramineous to medium brown, midrib region paler, ovate, 3.5–6 × 2–3 mm, membranous to cartilaginous, apex entire, rounded to subacute. |
basal spikelets absent; never proliferous, ovoid, terete, 5–15 × 2.5–4 mm, apex acute; proximal scale empty, deciduous, clasping 1/2 of culm, similar to floral scales; subproximal scale often empty; floral scales spiraled, 10–30, 5–6 per mm of rachilla, pale orange-brown, midribs green, broadly ovate, 2.5–4 × 1.5–2.5 mm, central area and often flanks papery to cartilaginous (or membranous), midrib evident, apex broadly rounded to obtuse. |
Flowers | perianth bristles brown, equaling achene or tubercle, densely spinulose; anthers brown, 2–2.4 mm. |
perianth bristles 5–6, bright brown, stout, slightly shorter than to equaling tubercle; spinules rarely absent, dense, variably from divaricate to antrorse, very short, and blunt to acute, to often much longer and retrorse, sharply acute; stamens 2–3; anthers 0.5–1 mm; styles 3-fid. |
Achenes | often very variable within one plant, ovoid to obovoid or obpyriform, 1.5–2.5 × 1–1.2 mm, beak to 1 × 0.6 mm. |
medium brown, obovoid to obpyriform, compressed (often obscurely) trigonous, angles evident, 0.9–1.7 × 0.8–2 mm; apex often constricted proximal to tubercle, very coarsely cancellate at 10X, each face with 8–10 longitudinal rows of large depressions. |
Tubercles | when present pale to dark brown, pyramidal, to 0.5 × 0.3 mm. |
bone-white to pale orange-brown, often red-spotted, subpyramidal, cross section plano-convex, 0.9–1.7(–2.4) × 0.7–1.2–2(–2.2) mm, at least as high and wide as achene, spongy, apex rounded, often with an abaxial acute projection. |
Eleocharis rostellata |
Eleocharis tuberculosa |
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Phenology | Fruiting late spring in south, summer–fall in north. | Fruiting spring (in south)–fall. |
Habitat | Very wet calcareous or brackish fens, springs, shores | Wet soil, freshwater, ponds, lakeshores, streams, meadows, pine woods, grasslands, disturbed places, bogs |
Elevation | 50–2400 m (200–7900 ft) | 0–500 m (0–1600 ft) |
Distribution |
AZ; CA; CT; DE; FL; ID; IL; IN; KS; LA; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TX; UT; VA; WA; WI; WY; BC; NS; ON; Mexico; West Indies (Cuba, Haiti, Puerto Rico)
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AL; AR; FL; GA; LA; MA; MS; NC; NJ; SC; TN; TX; NS
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Discussion | Eleocharis rostellata is highly competitive, often forming large monospecific colonies. The South American E. platypus C. B. Clarke is often treated as a synonym of E. rostellata. Eleocharis rostellata superfically closely resembles E. suksdorfiana in its culms, spikelets, and achenes, but differs in the absence of creeping rhizomes, presence of stoloniferous culms, absence of a flower in the proximal scale, and achene surface details. The collection of E. rostellata I have seen from Miami-Dade County, Florida, is from 1877. I have not seen vouchers for Archuleta County, Colorado, by H. D. Harrington (1954), or for the localities in Montana and South Carolina, which are based on the map in H. K. Svenson (1934). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis tuberculosa evidently is closely related to E. tortilis, from which it differs in its larger tubercles, culms that are always elliptic in cross section, and perianth bristle spinules that are often divaricate to antrorse and often blunt. The specimens of E. tuberculosa from throughout its range which are like E. tortilis in having retrorsely spinulose perianth bristles have been named E. tuberculosa forma retrorsa Svenson (type from Massachusetts). Plants from Nova Scotia with nearly smooth bristles have been named E. tuberculosa var. pubnicoensis Fernald and E. tuberculosa forma pubnicoensis (Fernald) Svenson. We have not seen voucher specimens for literature reports of Eleocharis tuberculosa from Connecticut, Delaware, Maine, Maryland, New Hampshire, New York, Pennsylvania, Rhode Island, or Virginia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 90. | FNA vol. 23, p. 92. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Rostellatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus rostellatus | Scirpus tuberculosus, E. tuberculosa var. pubnicoensis |
Name authority | (Torrey) Torrey: Fl. New York 2: 347. (1843) | (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 152. (1817) |
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