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beak spike-rush, beak spikesedge, walking sedge, walking spikerush

Baldwin's spike-rush

Habit Plants densely tufted, mat-forming by means of rooting culm tips. Plants annual(?), often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent.
Culms

1.5–3 times as wide as thick, 20–100 cm × 0.35–2 mm, firm to hard, wiry, with to 8 subacute ribs, rarely nearly smooth;

some culms arching or decumbent and rooting at tips.

often arching, sometimes spotted or streaked red-brown, quadrangular or broadly elliptic, sulcate or not, 0.5–25(–30) cm × 0.1–0.4 mm, soft.

Leaves

distal leaf sheaths not splitting abaxially, proximally dark red to brown, apex usually reddish.

distal leaf sheaths persistent or disintegrating, stramineous, green, red-brown, or spotted or streaked red-brown, membranous, translucent, apex obtuse or acute.

Spikelets

ovoid, 5–17 × 2.5–5 mm, apex acute;

spikelets on stolons rudimentary, non-flowering, proliferous when rooting;

proximal scale empty, amplexicaulous, ovate, 2–4 mm;

subproximal scale with flower;

floral scales 20–40, 2–3 per mm of rachilla, stramineous to medium brown, midrib region paler, ovate, 3.5–6 × 2–3 mm, membranous to cartilaginous, apex entire, rounded to subacute.

basal spikelets usually present, pistillate; often proliferous, ovoid, ellipsoid, or linear, clearly laterally compressed, 2.3–6.9 × 0.5–2 mm, apex acute, proximal scale empty, amplexicaulous, similar to floral scales (midribs sometimes more prominent and sometimes slightly prolonged beyond lamina);

subproximal scale with a flower;

floral scales clearly distichous, 3–9, 3–4 per mm of rachilla, yellowish, pale brown, or red-brown, spotted, streaked, or mottled red-brown, narrowly ovate to oblong, (1.5–)2–5 × 0.4–0.6(–1.2) mm, membranous, apex rounded, obtuse or rarely acute, midribs keeled.

Flowers

perianth bristles brown, equaling achene or tubercle, densely spinulose;

anthers brown, 2–2.4 mm.

perianth bristles 3–7, incurved, colorless, white, or red-brown, shorter than or equaling achene;

spinules sparse, retrorse, sharply acute;

stamens 3;

anthers 1.4–2.2 mm;

styles 3-fid.

Achenes

often very variable within one plant, ovoid to obovoid or obpyriform, 1.5–2.5 × 1–1.2 mm, beak to 1 × 0.6 mm.

whitish or greenish, spotted dark green, obovoid, trigonous, angles prominent, 0.6–0.9 × 0.4–0.6 mm, apex constricted proximal to tubercle, smooth to finely cancellate at 10–20X.

Tubercles

when present pale to dark brown, pyramidal, to 0.5 × 0.3 mm.

pale to red-brown, pyramidal, trigonous, 0.2–0.3(–0.4) × 0.2–0.5 mm.

Eleocharis rostellata

Eleocharis baldwinii

Phenology Fruiting late spring in south, summer–fall in north. Fruiting early spring–winter.
Habitat Very wet calcareous or brackish fens, springs, shores Freshwater to moist terrestrial sites, pine savannas, cypress ponds, lakeshores
Elevation 50–2400 m (200–7900 ft) 0–60 m (0–200 ft)
Distribution
from FNA
AZ; CA; CT; DE; FL; ID; IL; IN; KS; LA; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TX; UT; VA; WA; WI; WY; BC; NS; ON; Mexico; West Indies (Cuba, Haiti, Puerto Rico)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; NC; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis rostellata is highly competitive, often forming large monospecific colonies. The South American E. platypus C. B. Clarke is often treated as a synonym of E. rostellata. Eleocharis rostellata superfically closely resembles E. suksdorfiana in its culms, spikelets, and achenes, but differs in the absence of creeping rhizomes, presence of stoloniferous culms, absence of a flower in the proximal scale, and achene surface details. The collection of E. rostellata I have seen from Miami-Dade County, Florida, is from 1877. I have not seen vouchers for Archuleta County, Colorado, by H. D. Harrington (1954), or for the localities in Montana and South Carolina, which are based on the map in H. K. Svenson (1934).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Coarse plants of Eleocharis baldwinii usually bear more achenes per spikelet, dark colored achenes, culms gray-green when dried, and yellowish sheaths. Finer plants often bear few achenes per spikelet, whitish achenes, culms that are mid green when dried, and reddish sheaths. Achene color may relate to maturity, with darker achenes being fully mature. Although these variants appear to intergrade, they may be recognizable taxonomically. The longer floral scales in the spikelet are usually 3.5– 5 mm.

We have not seen vouchers to verify the report of Eleocharis baldwinii from Virginia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 90. FNA vol. 23, p. 98.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Rostellatae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus rostellatus Chaetocyperus baldwinii
Name authority (Torrey) Torrey: Fl. New York 2: 347. (1843) (Torrey) Chapman: Fl. South. U.S., 519. (1860)
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