Eleocharis rostellata |
Eleocharis ambigens |
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beak spike-rush, beak spikesedge, walking sedge, walking spikerush |
ambiguous spikesedge, éléocharide ambiquë |
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Habit | Plants densely tufted, mat-forming by means of rooting culm tips. | Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 1–3 cm, scales persistent, 5–10 mm, membranous, not fibrous. |
Culms | 1.5–3 times as wide as thick, 20–100 cm × 0.35–2 mm, firm to hard, wiry, with to 8 subacute ribs, rarely nearly smooth; some culms arching or decumbent and rooting at tips. |
terete, often with to 16 blunt ridges when dry, 25–80 cm × 0.5–1.5 mm, firm, internally spongy. |
Leaves | distal leaf sheaths not splitting abaxially, proximally dark red to brown, apex usually reddish. |
distal leaf sheaths persistent, not splitting, proximally dark red, distally red to stramineous or green, often callose, thinly papery (to membranous), apex dark red-brown, obtuse to subtruncate, tooth sometimes present on some culms, to 0.3 mm. |
Spikelets | ovoid, 5–17 × 2.5–5 mm, apex acute; spikelets on stolons rudimentary, non-flowering, proliferous when rooting; proximal scale empty, amplexicaulous, ovate, 2–4 mm; subproximal scale with flower; floral scales 20–40, 2–3 per mm of rachilla, stramineous to medium brown, midrib region paler, ovate, 3.5–6 × 2–3 mm, membranous to cartilaginous, apex entire, rounded to subacute. |
ovoid to lanceoloid, 5–23 × 2–3(–4) mm, apex acute (to obtuse); proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales often spreading in fruit, 10–60, 3–4 per mm of rachilla, medium brown, rarely red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, in distal part lanceolate, 2.5–3.5 × 1.7 mm, entire, apex acute to rarely obtuse, often carinate in distal part of spikelet. |
Flowers | perianth bristles brown, equaling achene or tubercle, densely spinulose; anthers brown, 2–2.4 mm. |
perianth bristles 2–4(–5), brown, slender to stout, usually very unequal, rudimentary to equaling achene; stamens 3; anthers brown, 1.5–2 mm; styles 2-fid, sometimes some 3-fid. |
Achenes | often very variable within one plant, ovoid to obovoid or obpyriform, 1.5–2.5 × 1–1.2 mm, beak to 1 × 0.6 mm. |
not persistent, dark yellow or stramineous, obovoid to obpyriform, all biconvex or some (very rarely all) compressedtrigonous, angles obscure, 1–1.5 × 0.9–1.25 mm, apex rounded, neck absent (to very short), finely rugulose at 10–20X, with 20 or more horizontal ridges in vertical series and/or finely cancellate at 10–30X. |
Tubercles | when present pale to dark brown, pyramidal, to 0.5 × 0.3 mm. |
brown or proximally whitish, pyramidal, depressed, rarely some as high as wide, 0.15–0.3(–0.5) × (0.3–)0.4–0.6 mm. |
2n | = 44, 45, 46. |
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Eleocharis rostellata |
Eleocharis ambigens |
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Phenology | Fruiting late spring in south, summer–fall in north. | Fruiting spring–summer. |
Habitat | Very wet calcareous or brackish fens, springs, shores | Coastal (rarely inland) fresh to brackish pond shores, marshes, disturbed places |
Elevation | 50–2400 m (200–7900 ft) | 0–10 m (0–0 ft) |
Distribution |
AZ; CA; CT; DE; FL; ID; IL; IN; KS; LA; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TX; UT; VA; WA; WI; WY; BC; NS; ON; Mexico; West Indies (Cuba, Haiti, Puerto Rico)
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CT; FL; GA; LA; MA; MS; NC; NJ; RI; SC; TX; VA; QC |
Discussion | Eleocharis rostellata is highly competitive, often forming large monospecific colonies. The South American E. platypus C. B. Clarke is often treated as a synonym of E. rostellata. Eleocharis rostellata superfically closely resembles E. suksdorfiana in its culms, spikelets, and achenes, but differs in the absence of creeping rhizomes, presence of stoloniferous culms, absence of a flower in the proximal scale, and achene surface details. The collection of E. rostellata I have seen from Miami-Dade County, Florida, is from 1877. I have not seen vouchers for Archuleta County, Colorado, by H. D. Harrington (1954), or for the localities in Montana and South Carolina, which are based on the map in H. K. Svenson (1934). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis ambigens is usually treated as a synonym of E. fallax because some specimens from Massachusetts to Georgia have some 3-fid styles and some obscurely trigonous achenes. A specimen of E. ambigens from Louisiana has trigonous achenes. Eleocharis ambigens is very similar to and perhaps better included in E. uniglumis (S.-O. Strandhede 1967). Intermediates with E. macrostachya occur in Louisiana and Texas. The depressed tubercles are diagnostic of E. ambigens. Eleocharis ambigens is sometimes confused with E. montevidensis, which is readily distinguished by its floral scales with broadly rounded apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 90. | FNA vol. 23, p. 78. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Rostellatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | Scirpus rostellatus | |
Name authority | (Torrey) Torrey: Fl. New York 2: 347. (1843) | Fernald: Rhodora 37: 394. (1935) |
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