Eleocharis robbinsii |
Eleocharis tortilis |
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Robbin's sppike-rush, Robbins' spikerush, Robbins' spikesedge, éléocharide Robbins |
twisted spikerush |
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Habit | Plants perennial; rhizomes (0.5–)1–2 mm thick, longer internodes 2–3 cm, scales 5–7 mm; tubers sometimes present, apical, ovoid, 4–8 × 3–4 mm. | Plants perennial, forming dense clumps, not stoloniferous, rhizomatous. |
Rhizomes | usually present, caudexlike, ascending, 1 mm thick, hidden by crowded culms; internodes and scales not evident. |
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Culms | acutely trigonous; spikelet-bearing culms 16–70 cm × 0.7–0.9 mm; when submersed plants often forming numerous, filiform, flaccid culms without spikelets, sometimes with whorls of slender branches, 0.1–0.3 mm wide; soft, sometimes septate-nodulose when aquatic, internally spongy, transverse septa incomplete. |
sometimes spirally twisted, usually minutely red-spotted at 30X, acutely triangular or sometimes elliptic, 15–70 cm × 0.3–0.5(–1) mm, soft (to firm), often minutely granular at 10X. |
Leaves | distal leaf sheaths persistent or decaying, membranous, apex obtuse to acuminate. |
distal leaf sheaths persistent, stramineous to green, often minutely red-spotted, membranous to papery, apex subacute to narrowly acute, often with toothlike callus. |
Spikelets | sometimes proliferous (when submerged), 9–33 × 1.5–3 mm; rachilla joints bearing prominent winglike remnants of floral scales; proximal scale with a flower, amplexicaulous, (5–)6–9.8 mm; floral scales 4–18, 0.5–1 per mm of rachilla, stramineous to pale brown, often minutely dotted reddish, without or rarely with darker submarginal band, narrowly ovate to lanceolate, 5–7.8 × 2–3 mm, thickly papery, membranous toward margins, apex narrowly rounded to acute. |
basal spikelets absent; never proliferous, ovoid to lanceoloid or ellipsoid, terete, 4–12 × 2–4 mm, apex acute; proximal scale empty, deciduous, clasping 1/2 of culm, similar to floral scales; subproximal scale often empty; floral scales spiraled, 10–25, 5–6 per mm of rachilla, pale orange-brown, midrib regions broadly pale green to colorless, broadly ovate, 2–2.5(–3.5) × 1.5–2 mm, central area and often flanks papery to cartilaginous or sometimes membranous, midrib evident, apex subacute (to rounded). |
Flowers | perianth bristles 6–7, stramineous to reddish brown, proximally slightly flattened, subequal to equal, much exceeding to rarely shorter than achene, 3–5 mm, retrorsely spinulose; anthers yellow to reddish, 1.6–3.2 mm; styles 3-fid. |
perianth bristles 4–6, bright brown, stout, equaling to slightly exceeding tubercle; spinules dense, retrorse, sharply acute; stamens 3; anthers 0.5–0.9 mm; styles 3-fid. |
Achenes | stramineous or medium brown, biconvex or compressed trigonous, narrowly obpyriform, 1.9–2.6 × 1–1.4 mm, adaxial face with 15–22 rows of rectangular, transversely elongated or nearly isodiametric cells, clearly sculptured at 10–15X, apex usually conspicuously constricted to short neck 0.4–0.7 mm wide, usually wider at tubercle base. |
medium brown or yellowish or greenish, obpyriform, obtusely (often obscurely) compressed trigonous, angles evident, 1.2–1.7(–2.4) × 0.9–1.2(–2.2) mm, apex constricted proximal to tubercle, coarsely cancellate at 10X, each face with 8–10 longitudinal rows of distinctly depressed, enlarged cells. |
Tubercles | stramineous to medium brown, high-pyramidal, 0.5–1.1 × 0.3–0.7 mm. |
brown to greenish (or bone-white), pyramidal, trigonous or cross section plano-convex, 0.3–0.7(–1) × 0.3–0.6(–0.9) mm, distinctly shorter or narrower than achene, apex acute, often acuminate. |
Eleocharis robbinsii |
Eleocharis tortilis |
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Phenology | Fruiting late spring–late fall. | Fruiting spring (in south)–fall. |
Habitat | Shallow waters of fresh lakes and ponds with sandy-peaty soils | Wet soil, freshwater, acidic places, seeps, bogs, ditches |
Elevation | 10–500 m (0–1600 ft) | 10–100 m (0–300 ft) |
Distribution |
AL; CT; DE; FL; GA; MA; ME; MI; MN; NC; NH; NJ; OH; SC; VA; WI; NB; NS; ON; QC
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AL; AR; FL; GA; LA; MD; MS; NC; NJ; SC; TN; TX; VA
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Discussion | I have not seen voucher specimens for literature reports of Eleocharis robbinsii from Indiana, Pennsylvania, or Rhode Island. Plants from South Carolina with the achene surface cells nearly isodiametric, the achene apex spongy, and the anthers to 3.2 mm may represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis tortilis differs consistently from E. tuberculosa only in tubercle size. Most specimens of E. tortilis have culms that are sharply trigonous and no more than 0.5 mm wide. However, about two-thirds of the specimens of E. tortilis from Texas resemble E. tuberculosa in having culms that are elliptic or subelliptic in cross section and that sometimes reach 1 mm wide. Those elliptic-culmed plants may deserve taxonomic recognition. I have not seen voucher specimens for reports in the literature (H. K. Svenson 1937; M. L. Brown and R. G. Brown 1984) for Alabama (Mobile, 1800s), Delaware (1908), Maryland (1863–1878), and New York (Long Island, 1903). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 117. | FNA vol. 23, p. 92. |
Parent taxa | Cyperaceae > Eleocharis > subg. Limnochloa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus tortilis | |
Name authority | Oakes: Mag. Hort. Bot. 7: 178. (1841) | (Link) Schultes: Mant. 2: 92. (1824) |
Web links |