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creeping spikerush, radical fiber optic spikerush, root spikerush

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

Habit Plants perennial, mat-forming; rhizomes 0.2–0.5 mm thick, longer internodes 5–10 mm, scales not evident. Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent.
Culms

obscurely to ca. 10-ribbed when dry, terete, 1–12 cm × 0.4–1 mm, very soft, spongy.

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

Leaves

distal leaf sheaths fugaceous, colorless, translucent, apex blunt.

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

Spikelets

ovoid, 2–3 × 1–1.5 mm, apex acute;

scales 5–15, 5–7 per mm of rachilla, colorless to stramineous, rarely slightly reddish, medially broadly greenish to colorless, faintly 3–5-veined, often ovate, membranous, fleshy;

proximal scale 2–2.5 × 0.7 mm, other scales 1.5–2 × 0.7 mm, midrib obscure to slightly keeled, apex acute to rounded.

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

Flowers

perianth bristles equaling achene;

anthers yellow to brown, 0.3–0.5 mm.

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

Achenes

brownish, angles plus longitudinal ridges ca. 7, narrowly obovoid, 2 times longer than wide, 0.75–0.9 × 0.35–0.4 mm, trabeculae ca. 50, distinct.

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

Tubercles

brown, pyramidal, 0.15 × 0.15–0.2 mm.

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

Eleocharis radicans

Eleocharis quadrangulata

Phenology Fruiting spring–winter (Apr–Dec). Fruiting early summer–winter.
Habitat Stream alluvium, lake margins, meadows, seeps, bogs Shallow water of fresh lake and pond shores, marshes
Elevation 100–1400 Ariz., Calif., Fla., La., Mich., Okla., Tex., Va. 10–600 m (0–2000 ft)
Distribution
from FNA
HI; n Mexico; West Indies; Central America (El Salvador); South America (Argentina, Chile)
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 112. FNA vol. 23, p. 120.
Parent taxa Cyperaceae > Eleocharis > subg. Scirpidium Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus radicans, E. acicularis var. radicans, E. lindheimeri Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus
Name authority (Poiret) Kunth: Enum. Pl. 2: 142. (1837) (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817)
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