Eleocharis quinqueflora
Eleocharis kamtschatica
few-flower spike-rush, few-flower spikesedge, éléocharide à cinq fleurs
Kamchatka spike-rush, éléocharide du kamtchatka
erect, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times wider than thick, 5–35 cm × 0.2–0.5(–1.2) mm, soft to hard;
culm tufts often proximally bulbous (if bulbous then tunicated by papery-fibrous scales).
terete, often with to 10 blunt ridges when dry, 4–60 cm × 0.3–1.8 mm, soft to firm, internally spongy.
distal leaf sheaths stramineous to brown or reddish proximally, green to stramineous or brown distally, membranous to papery, apex often reddish, subtruncate to acute.
distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally red or stramineous or green, not callose, membranous, apex usually pale red, broadly obtuse to narrowly acuminate, tooth absent.
3–8 × 1.5–4 mm;
proximal scale with a flower, seldom empty, 2–5 mm, 1/2 or more as long as spikelet;
floral scales 3–10 per spikelet, 2.5–6 × 1.5–2.5 mm.
ovoid to lanceoloid, 4–20 × 3–6 mm, apex acute;
proximal scale amplexicaulous, entire;
subproximal scale with flower;
floral scales often spreading in fruit, 5–20, 1–2 per mm of rachilla, entirely blackish brown to red-brown, or midrib regions paler, broadly ovate, 2.5–5 × 2–2.5 mm, apex rounded to subacute, not carinate.
perianth bristles (0–)3–6, often unequal, rudimentary to equaling tubercle, stout to slender, spinules dense to apparently absent;
anthers 1.5–2.7(–3.5) mm.
perianth bristles 0–6(–9), light brown to stramineous, slender to stout, often unequal, much shorter than to equaling achene;
stamens 3;
anthers yellow-brown to medium brown, 1.3–2.5 mm;
styles 2-fid.
stramineous to medium brown or gray-brown, equilaterally trigonous to compressed-trigonous, rarely some biconvex, obpyriform (to obovoid), 1.6–2.3 × 0.7–1.3 mm, beak variable.
not persistent, yellow, stramineous, or medium brown, obovoid (mostly very broadly so), thickly biconvex, angles obscure, 1–1.5(–1.9) × 1–1.6 mm, apex truncate, neck absent or very short, finely reticulate at 20–30X, or rugulose with 20 or more horizontal ridges in a vertical series, or sometimes smooth.
rarely absent, 0.3–0.4 × 0.2–0.3 mm.
whitish, outline subrectangular or trapezoidal, sometimes broadly rounded, seldom pyramidal, (0.5–) 0.7–1.5 × (0.5–)0.8–1.4 mm, usually with vertical rows of depressions (cancellate).
= 12.
Eleocharis quinqueflora
Eleocharis kamtschatica
The chromosome numbers for Eleocharis quinqueflora reported for North America (2n = 80) are in doubt because vouchers and other information are lacking. The often-cited n = 10 is probably erroneous. S.-O. Strandhede and R. M. T. Dahlgren (1968) gave 2n = 132 and 134 from Scandinavia. Recognition of infraspecific taxa within E. quinqueflora is premature pending a worldwide revision of subg. Zinserlingia. It has been reported from North Dakota, although I have not seen specimens. About five varieties and subspecies of E. quinqueflora have been described worldwide.
Most specimens from eastern North America and some from the West can be placed in Eleocharis quinqueflora subsp. fernaldii (Svenson) Hultén, which is characterized by its small size (culms to 15 cm × 0.5 mm) and small bulbs. Specimens of E. quinqueflora from 2000–3600 m in California, which are atypical, especially in that the proximal scales of the spikelets do not subtend flowers, may deserve taxonomic recognition. Those plants are also small, with culms only to 15 cm × 0.5 mm; hard caudices are often present at the culm-tuft bases; small, narrowly ovoid bulbs are sometimes present; and perianth bristles are absent or rudimentary. Very few specimens of E. quinqueflora are intermediate with E. suksdorfiana.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The markedly different chromosome counts reported from North America (2n = 12, from the Queen Charlotte Islands, British Columbia) and from Asia (2n = 38–40, 42, 56) suggest that more than one taxon should be recognized within Eleocharis kamtschatica as currently broadly defined. The three similar species E. kamtschatica, E. komarovii, and E. sachalinensis have been recognized for Russia (I. D. Zinserling 1935); more recent authors have recognized only E. kamtschatica (T. V. Egorova 1981 and A. E. Kozhevnikov 1988, for the former U.S.S.R.; J. Ohwi 1965, for Japan; S.-O Strandhede 1967; E. Hultén 1968 for Alaska; Tang T. and Wang F. T. 1961, for China). Pending a taxonomic revision, the following five names, all from eastern Asia, should probably be treated as synonyms of E. kamtschatica: E. komarovii Zinserling, E. mitrata (Franchet & Savatier ex Makino, E. sachalinensis (Meinschauser) B. Fedtschenko, E. savatieri Svenson, and E. triflora Komarov, not E. triflora Boeckeler. In E. kamtschatica in the broad sense, the size and form of the achenes and tubercles, on which most of the probable synonyms listed above are based, and the form of the distal leaf sheath summits do not seem to be correlated with each other or with other characters. In North America E. kamtschatica is known only from coastal habitats; in Asia it also occurs inland, especially around hot springs.
North American plants of Eleocharis kamtschatica seem to be separable into two groups: (1) Smaller plants, with culms mostly shorter than 20 cm, spikelets 5–13 × 3–4 mm, and floral scales 2.5–4 mm are known from about 20 localities on the Pacific Coast from Kodiak Island, Alaska, to the Queen Charlotte Islands and Prince Rupert, British Columbia, as well as from Hudson Bay in Manitoba, James Bay and the Gulf of St. Lawrence in Quebec, and Labrador. These small plants include the voucher specimens of the 2n = 12 chromosome count from British Columbia, and seem to fit E. kamtschatica as described by I. D. Zinserling (1935). (2) More robust plants, with culms mostly 20–70 cm, spikelets 7–20 × 4–6 mm, and floral scales 4–5 mm, are known from about 12 localities from Norton Sound in west-central Alaska to central British Columbia. These robust plants fit the descriptions of E. sachalinensis by I. D. Zinserling(1935), of E. savatieri Svenson from Japan (H. K. Svenson 1939), and of E. kamtschatica as described by J. Ohwi (1965), and are very similar to several specimens from Japan that I have seen.
Because Eleocharis kamtschatica closely resembles E. uniglumis except for size of the tubercles, plants without achenes cannot be reliably identified to species. The floral scales and receptacles in E. kamtschatica are very sparsely placed, with only 1–2 scales per mm of rachilla, compared with 3–4 mm of rachilla in most North American plants of E. uniglumis. Some North American collections, e.g., from the Kobuk sand dunes in northwestern Alaska, as E. uniglumis, and James Bay in Quebec, as E. kamtschatica, are intermediate between E. kamtschatica and E. uniglumis in the relative sizes of the tubercles and achenes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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