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few-flower spike-rush, few-flower spikesedge, éléocharide à cinq fleurs

bald spike-rush, red-footed spikesedge, red-stem spikerush, redfoot spike-rush, spikesedge, éléocharide à tiges rouges

Habit Plants perennial; rhizomes 0.2–1 mm thick, scales persistent or fugaceous, 2–4(–7) mm, thinly membranous, not fibrous; resting buds often present on rhizomes or among culm bases, broadly to narrowly ovoid, 3–6(–10) × 2–5 mm; caudices absent, rarely present, soft or rarely hard, 0.5 mm thick. Plants perennial, mat-forming; rhizomes evident, long, 0.5–1.5 mm thick, soft to firm, cortex often breaking loose, longer internodes 5–30 mm, scales often fugaceous, 4–8 mm, membranous, not fibrous.
Culms

erect, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times wider than thick, 5–35 cm × 0.2–0.5(–1.2) mm, soft to hard;

culm tufts often proximally bulbous (if bulbous then tunicated by papery-fibrous scales).

terete, often with 8–12 blunt ridges when dry, 8–80 cm × 0.3–1.4 mm, firm to soft, internally spongy.

Leaves

distal leaf sheaths stramineous to brown or reddish proximally, green to stramineous or brown distally, membranous to papery, apex often reddish, subtruncate to acute.

distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally green to stramineous, usually inflated, often callose, membranous to papery, apex often red-brown, broadly obtuse to subacute, tooth sometimes present, to 0.1 mm.

Spikelets

3–8 × 1.5–4 mm;

proximal scale with a flower, seldom empty, 2–5 mm, 1/2 or more as long as spikelet;

floral scales 3–10 per spikelet, 2.5–6 × 1.5–2.5 mm.

ovoid to lanceoloid or nearly cylindric, 3–18 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 15–50, 4–5 per mm of rachilla, medium brown to sometimes red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, apex rounded, in distal part lanceolate, apex entire, acute, 2–3.5 × 1.5–1.7 mm, mostly carinate.

Flowers

perianth bristles (0–)3–6, often unequal, rudimentary to equaling tubercle, stout to slender, spinules dense to apparently absent;

anthers 1.5–2.7(–3.5) mm.

perianth bristles 4 or absent, light brown to stramineous, stout, usually equal, equaling achene to slightly exceeding tubercle;

stamens 3;

anthers dark yellow to stramineous, 1–1.8 mm, apiculate;

styles 2-fid.

Achenes

stramineous to medium brown or gray-brown, equilaterally trigonous to compressed-trigonous, rarely some biconvex, obpyriform (to obovoid), 1.6–2.3 × 0.7–1.3 mm, beak variable.

not persistent, dark yellow, stramineous, or dark brown, obovoid to obpyriform, biconvex, angles obscure, 0.9–1.6 × 0.7–1.2 mm, apex rounded, neck absent to long, smooth at 30X, or sometimes finely rugulose at 10–30X with 20 or more horizontal ridges in a vertical series.

Tubercles

rarely absent, 0.3–0.4 × 0.2–0.3 mm.

brown to whitish, pyramidal, much higher than wide to, lower than wide, 0.35–0.65 × 0.2–0.6 mm.

2n

= 16, 18, 19, 20.

Eleocharis quinqueflora

Eleocharis erythropoda

Phenology Fruiting (spring–)summer. Fruiting summer.
Habitat Fens, wet meadows, seeps, springs, hot springs Non-calcareous or calcareous fresh or brackish shores, marshes, meadows, fens, disturbed places
Elevation 0–3600 m (0–11800 ft) 0–2300 m (0–7500 ft)
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; IL; IN; MA; ME; MI; MN; MT; NE; NH; NJ; NM; NV; NY; OH; OR; PA; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
[WildflowerSearch map]
Discussion

The chromosome numbers for Eleocharis quinqueflora reported for North America (2n = 80) are in doubt because vouchers and other information are lacking. The often-cited n = 10 is probably erroneous. S.-O. Strandhede and R. M. T. Dahlgren (1968) gave 2n = 132 and 134 from Scandinavia. Recognition of infraspecific taxa within E. quinqueflora is premature pending a worldwide revision of subg. Zinserlingia. It has been reported from North Dakota, although I have not seen specimens. About five varieties and subspecies of E. quinqueflora have been described worldwide.

Most specimens from eastern North America and some from the West can be placed in Eleocharis quinqueflora subsp. fernaldii (Svenson) Hultén, which is characterized by its small size (culms to 15 cm × 0.5 mm) and small bulbs. Specimens of E. quinqueflora from 2000–3600 m in California, which are atypical, especially in that the proximal scales of the spikelets do not subtend flowers, may deserve taxonomic recognition. Those plants are also small, with culms only to 15 cm × 0.5 mm; hard caudices are often present at the culm-tuft bases; small, narrowly ovoid bulbs are sometimes present; and perianth bristles are absent or rudimentary. Very few specimens of E. quinqueflora are intermediate with E. suksdorfiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis erythropoda is extremely variable. Intermediates with E. palustris variant b are common in the region of sympatry in the East, and with E. macrostachya variant b in the West, and E. erythropoda is sometimes difficult to distinguish from E. uniglumis.

Eleocharis calva Torrey is an invalid name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 114. FNA vol. 23, p. 75.
Parent taxa Cyperaceae > Eleocharis > subg. Zinserlingia Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus quinqueflorus, E. fernaldii, E. pauciflora, E. pauciflora var. fernaldii, E. quinqueflora subsp. fernaldii Scirpus glaucus
Name authority (Hartmann) O. Schwarz: Mitt. Thüring. Bot. Ges. 1: 89. (1949) Steudel: Syn. Pl. Glumac. 2: 76. (1855)
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