Eleocharis quadrangulata |
Eleocharis radicans |
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four-angle spikerush, square-stem spike-rush, square-stem spikesedge |
creeping spikerush, radical fiber optic spikerush, root spikerush |
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Habit | Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. | Plants perennial, mat-forming; rhizomes 0.2–0.5 mm thick, longer internodes 5–10 mm, scales not evident. |
Culms | acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete; plants never forming filiform, flaccid culms. |
obscurely to ca. 10-ribbed when dry, terete, 1–12 cm × 0.4–1 mm, very soft, spongy. |
Leaves | distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm. |
distal leaf sheaths fugaceous, colorless, translucent, apex blunt. |
Spikelets | not proliferous, (15–)20–76 × 3–5(–6) mm; rachilla joints bearing obscure winglike remnants of floral scales; proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm; floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse. |
ovoid, 2–3 × 1–1.5 mm, apex acute; scales 5–15, 5–7 per mm of rachilla, colorless to stramineous, rarely slightly reddish, medially broadly greenish to colorless, faintly 3–5-veined, often ovate, membranous, fleshy; proximal scale 2–2.5 × 0.7 mm, other scales 1.5–2 × 0.7 mm, midrib obscure to slightly keeled, apex acute to rounded. |
Flowers | perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth; anthers stramineous to red-brown, 2.3–2.9 mm; styles 3-fid, sometimes 2-fid. |
perianth bristles equaling achene; anthers yellow to brown, 0.3–0.5 mm. |
Achenes | yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide. |
brownish, angles plus longitudinal ridges ca. 7, narrowly obovoid, 2 times longer than wide, 0.75–0.9 × 0.35–0.4 mm, trabeculae ca. 50, distinct. |
Tubercles | dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy. |
brown, pyramidal, 0.15 × 0.15–0.2 mm. |
Eleocharis quadrangulata |
Eleocharis radicans |
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Phenology | Fruiting early summer–winter. | Fruiting spring–winter (Apr–Dec). |
Habitat | Shallow water of fresh lake and pond shores, marshes | Stream alluvium, lake margins, meadows, seeps, bogs |
Elevation | 10–600 m (0–2000 ft) | 100–1400 Ariz., Calif., Fla., La., Mich., Okla., Tex., Va. |
Distribution |
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
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HI; n Mexico; West Indies; Central America (El Salvador); South America (Argentina, Chile)
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Discussion | We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee. The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 23, p. 120. | FNA vol. 23, p. 112. |
Parent taxa | Cyperaceae > Eleocharis > subg. Limnochloa | Cyperaceae > Eleocharis > subg. Scirpidium |
Sibling taxa | ||
Synonyms | Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus | Scirpus radicans, E. acicularis var. radicans, E. lindheimeri |
Name authority | (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) | (Poiret) Kunth: Enum. Pl. 2: 142. (1837) |
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