The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

creeping spikerush, radical fiber optic spikerush, root spikerush

Habit Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. Plants perennial, mat-forming; rhizomes 0.2–0.5 mm thick, longer internodes 5–10 mm, scales not evident.
Culms

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

obscurely to ca. 10-ribbed when dry, terete, 1–12 cm × 0.4–1 mm, very soft, spongy.

Leaves

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

distal leaf sheaths fugaceous, colorless, translucent, apex blunt.

Spikelets

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

ovoid, 2–3 × 1–1.5 mm, apex acute;

scales 5–15, 5–7 per mm of rachilla, colorless to stramineous, rarely slightly reddish, medially broadly greenish to colorless, faintly 3–5-veined, often ovate, membranous, fleshy;

proximal scale 2–2.5 × 0.7 mm, other scales 1.5–2 × 0.7 mm, midrib obscure to slightly keeled, apex acute to rounded.

Flowers

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

perianth bristles equaling achene;

anthers yellow to brown, 0.3–0.5 mm.

Achenes

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

brownish, angles plus longitudinal ridges ca. 7, narrowly obovoid, 2 times longer than wide, 0.75–0.9 × 0.35–0.4 mm, trabeculae ca. 50, distinct.

Tubercles

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

brown, pyramidal, 0.15 × 0.15–0.2 mm.

Eleocharis quadrangulata

Eleocharis radicans

Phenology Fruiting early summer–winter. Fruiting spring–winter (Apr–Dec).
Habitat Shallow water of fresh lake and pond shores, marshes Stream alluvium, lake margins, meadows, seeps, bogs
Elevation 10–600 m (0–2000 ft) 100–1400 Ariz., Calif., Fla., La., Mich., Okla., Tex., Va.
Distribution
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
HI; n Mexico; West Indies; Central America (El Salvador); South America (Argentina, Chile)
[WildflowerSearch map]
[BONAP county map]
Discussion

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 120. FNA vol. 23, p. 112.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Scirpidium
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus Scirpus radicans, E. acicularis var. radicans, E. lindheimeri
Name authority (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) (Poiret) Kunth: Enum. Pl. 2: 142. (1837)
Web links