The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine

Habit Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent.
Culms

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

Leaves

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

Spikelets

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

2–4 × 1–2 mm, sometimes absent in deeper water;

proximal scale 1/2 or more of spikelet length;

floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute.

Flowers

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose;

anthers 0.7–1.2 mm.

Achenes

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X.

Tubercles

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

0.1–0.2 × 0.15 mm.

Tubers

terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm;

tubers among culm bases straight, narrowly fusiform, 4–5 mm.

2n

= 10 (Europe).

Eleocharis quadrangulata

Eleocharis parvula

Phenology Fruiting early summer–winter. Fruiting summer–fall (north) or late winter–fall (far south).
Habitat Shallow water of fresh lake and pond shores, marshes Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches
Elevation 10–600 m (0–2000 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 120. FNA vol. 23, p. 106.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus Scirpus parvulus, E. pygmaea, S. nanus
Name authority (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836)
Web links