The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

decumbent spikerush

Habit Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. Plants perennial, densely tufted; rhizomes often hidden by culms and roots, fairly long, 3–4 mm thick, hard, cortex persistent, longer internodes from very short to 5 mm, scales usually clearly evident, disintegrating to fibers, 20–25 mm, papery.
Culms

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

terete, often with 10–18 blunt ridges when dry, 10–50 cm × 0.3–2 mm, firm to rigid, spongy.

Leaves

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

distal leaf sheaths persistent, not splitting, proximally brown or reddish, distally stramineous, brown, reddish or green, often inflated, papery, apex mostly dark red-brown, subtruncate to obtuse, often callose, tooth absent.

Spikelets

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

ovoid, 3–8 × 2–2.5 mm, apex acute;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 10–20, 3 per mm of rachilla, orange-brown, midrib regions often greenish, ovate, 3–3.5 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

Flowers

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

perianth bristles 6, stramineous, stout, nearly equal, mostly equaling or exceeding tubercle, (0.5–)1–2.2 mm, prominently retrorsely spinulose;

stamens 3(?);

anthers dark yellow to stramineous, 1.2–1.5;

styles 3-fid.

Achenes

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

falling with scales, dark brown, obpyriform, nearly eqilaterally- to greatly compressed-trigonous, angles slightly prominent, 1–1.3 × 0.75–0.9 mm, neck absent or short, finely rugulose at 20X with more than 20 horizontal ridges in vertical series, or reticulate or cancellate at 20–30X.

Tubercles

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

well developed, whitish, pyramidal, as high as wide to much lower than wide, 0.2–0.6 × 0.4–0.7 mm.

Eleocharis quadrangulata

Eleocharis decumbens

Phenology Fruiting early summer–winter. Fruiting summer–fall.
Habitat Shallow water of fresh lake and pond shores, marshes Wet fresh meadows, seeps, and lakeshores, in interior montane conifer forests and alpine zones
Elevation 10–600 m (0–2000 ft) 700–3500 m (2300–11500 ft)
Distribution
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis decumbens is known only from Shasta and Tulare counties in California.

Although Eleocharis decumbens has long been ignored or treated as a variety of E. montevidensis, it clearly is a very distinct species, from which it differs especially in its thick rhizomes with fibrous scales and its acute floral scales.

Specimens of Eleocharis decumbens without rhizomes or achenes are easily confused with the apparently closely related E. bolanderi, which often may be distinguished by culms no more than 0.5 mm wide and spikelets with scales no more than 3 mm long. The tubercles of E. decumbens are usually well differentiated from the achenes and about as high as wide; the tubercles of E. bolanderi are often poorly developed and much lower than wide. Three collections from Jackson and Klamath counties in Oregon, lack achenes but are probably E. decumbens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 120. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus E. montevidensis var. decumbens
Name authority (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) C. B. Clarke: Bull. Misc. Inform., addit. ser. 8: 23. (1908)
Web links