The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

glades spikerush

Habit Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. Plants perennial, densely cespitose; rhizomes concealed by persistent culm bases, short, 4–5 mm thick, hard, cortex persistent, internodes crowded, scales decaying to coarse fibers, 1 cm, papery.
Culms

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

greatly compressed, 4–10 times wider than thick, often with 1 or 2 sharp ridges on 1 side, (8–)20–35 cm × 0.7–2.3 mm, hard, finely striate, spongy.

Leaves

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

distal leaf sheaths persistent, not splitting, proximally red or stramineous, distally green to stramineous, inflated, papery, apex dark brown, broadly obtuse to subtruncate, callose, tooth absent.

Spikelets

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

ovoid, 4–9 × 2.5–4 mm, apex acute;

proximal scale clasping 2/3–3/4 of culm, apex 2-fid;

subproximal scale empty or with a flower;

floral scales spreading in fruit, 30–60, 6–9 per mm of rachilla, medium or pale brown, midrib region often paler, ovate-lanceolate, 2.5–3.5 × 1.5 mm, apex 2-fid, carinate in distal part of spikelet.

Flowers

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

perianth bristles 0(–5), stramineous to pale brown, 1/2 of to equaling achene length;

stamens 3;

anthers orange-brown, 0.7–1.5 mm;

styles 3-fid.

Achenes

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

falling with scales, yellow-brown to medium brown, obovoid to obpyriform, nearly equilaterally trigonous, cross section circular to slightly compressed, angles obscure or evident, 0.9–1.1 × 0.6–0.75, neck usually very short, finely rugulose at 10–30X, 20 or more low, blunt horizontal ridges in vertical series.

Tubercles

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

brown, depressed-pyramidal, often rudimentary, 0.1–0.25 × 0.2–0.3 mm.

Eleocharis quadrangulata

Eleocharis bifida

Phenology Fruiting early summer–winter. Fruiting spring–summer (May–Jun).
Habitat Shallow water of fresh lake and pond shores, marshes Seasonally wet places on limestone, cedar (juniper) glades, stream beds, prairies, ditches
Elevation 10–600 m (0–2000 ft) 200–300 m (700–1000 ft)
Distribution
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; KY; TN
[BONAP county map]
Discussion

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis bifida was previously included in E. compressa. A very few specimens of E. compressa var. compressa from the states where E. bifida is known are like E. bifida except for their entire proximal floral scales and evident rhizomes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 120. FNA vol. 23, p. 85.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus
Name authority (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) S. G. Smith: Novon 11: 243, figs. 1F–J, 2. (2001)
Web links