Eleocharis quadrangulata |
Eleocharis baldwinii |
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four-angle spikerush, square-stem spike-rush, square-stem spikesedge |
Baldwin's spike-rush |
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Habit | Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. | Plants annual(?), often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. |
Culms | acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete; plants never forming filiform, flaccid culms. |
often arching, sometimes spotted or streaked red-brown, quadrangular or broadly elliptic, sulcate or not, 0.5–25(–30) cm × 0.1–0.4 mm, soft. |
Leaves | distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm. |
distal leaf sheaths persistent or disintegrating, stramineous, green, red-brown, or spotted or streaked red-brown, membranous, translucent, apex obtuse or acute. |
Spikelets | not proliferous, (15–)20–76 × 3–5(–6) mm; rachilla joints bearing obscure winglike remnants of floral scales; proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm; floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse. |
basal spikelets usually present, pistillate; often proliferous, ovoid, ellipsoid, or linear, clearly laterally compressed, 2.3–6.9 × 0.5–2 mm, apex acute, proximal scale empty, amplexicaulous, similar to floral scales (midribs sometimes more prominent and sometimes slightly prolonged beyond lamina); subproximal scale with a flower; floral scales clearly distichous, 3–9, 3–4 per mm of rachilla, yellowish, pale brown, or red-brown, spotted, streaked, or mottled red-brown, narrowly ovate to oblong, (1.5–)2–5 × 0.4–0.6(–1.2) mm, membranous, apex rounded, obtuse or rarely acute, midribs keeled. |
Flowers | perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth; anthers stramineous to red-brown, 2.3–2.9 mm; styles 3-fid, sometimes 2-fid. |
perianth bristles 3–7, incurved, colorless, white, or red-brown, shorter than or equaling achene; spinules sparse, retrorse, sharply acute; stamens 3; anthers 1.4–2.2 mm; styles 3-fid. |
Achenes | yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide. |
whitish or greenish, spotted dark green, obovoid, trigonous, angles prominent, 0.6–0.9 × 0.4–0.6 mm, apex constricted proximal to tubercle, smooth to finely cancellate at 10–20X. |
Tubercles | dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy. |
pale to red-brown, pyramidal, trigonous, 0.2–0.3(–0.4) × 0.2–0.5 mm. |
Eleocharis quadrangulata |
Eleocharis baldwinii |
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Phenology | Fruiting early summer–winter. | Fruiting early spring–winter. |
Habitat | Shallow water of fresh lake and pond shores, marshes | Freshwater to moist terrestrial sites, pine savannas, cypress ponds, lakeshores |
Elevation | 10–600 m (0–2000 ft) | 0–60 m (0–200 ft) |
Distribution |
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
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AL; AR; FL; GA; LA; MS; NC; SC; TX
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Discussion | We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee. The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Coarse plants of Eleocharis baldwinii usually bear more achenes per spikelet, dark colored achenes, culms gray-green when dried, and yellowish sheaths. Finer plants often bear few achenes per spikelet, whitish achenes, culms that are mid green when dried, and reddish sheaths. Achene color may relate to maturity, with darker achenes being fully mature. Although these variants appear to intergrade, they may be recognizable taxonomically. The longer floral scales in the spikelet are usually 3.5– 5 mm. We have not seen vouchers to verify the report of Eleocharis baldwinii from Virginia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 120. | FNA vol. 23, p. 98. |
Parent taxa | Cyperaceae > Eleocharis > subg. Limnochloa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus | Chaetocyperus baldwinii |
Name authority | (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) | (Torrey) Chapman: Fl. South. U.S., 519. (1860) |
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