Eleocharis parvula |
Eleocharis torticulmis |
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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine |
California twisted spikerush, twist-stem spikerush |
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Habit | Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick. | |
Culms | erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm; culm tufts not proximally bulbous. |
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Leaves | distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse. |
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Spikelets | 2–4 × 1–2 mm, sometimes absent in deeper water; proximal scale 1/2 or more of spikelet length; floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute. |
6–8 × 2–3 mm; proximal scale empty, 3–4 mm, shorter than to equaling spikelet; floral scales 8–10 per spikelet, 3.5–5 × 2 mm. |
Flowers | perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose; anthers 0.7–1.2 mm. |
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Perianth | bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose. |
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Achenes | stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X. |
stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm; beak 0.3–0.6 mm. |
Tubers | terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm; tubers among culm bases straight, narrowly fusiform, 4–5 mm. |
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Tubercles | 0.1–0.2 × 0.15 mm. |
0.25–0.6 × 0.3–0.55 mm. |
Anthers | 1.8 –3 mm. |
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2n | = 10 (Europe). |
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Eleocharis parvula |
Eleocharis torticulmis |
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Phenology | Fruiting summer–fall (north) or late winter–fall (far south). | Fruiting summer (Jun–Jul). |
Habitat | Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches | Fens, wet meadows, vernal ponds |
Elevation | 0–600 m (0–2000 ft) | 1100 m (3600 ft) |
Distribution |
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
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CA |
Discussion | Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 106. | FNA vol. 23, p. 115. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae | Cyperaceae > Eleocharis > subg. Zinserlingia |
Sibling taxa | ||
Synonyms | Scirpus parvulus, E. pygmaea, S. nanus | |
Name authority | (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) | S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001) |
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