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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine

Suksdorf spikerush, Suksdorf's spike-rush

Habit Plants perennial; rhizomes 0.5–1.5 mm thick, scales persistent, 5–10 mm, thinly membranous, not fibrous; resting buds absent (non-resting buds on rhizome apex ellipsoid, 10 × 2–5 mm); caudices present, hard, 2 mm thick.
Culms

erect, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times as wide as thick, (5–)10–40 cm × 0.5–1.2 mm, firm to hard, finely many-ridged at 10–20X;

culm tufts not proximally bulbous.

Leaves

distal leaf sheaths stramineous to brown or reddish proximally, green to stramineous distally, subtruncate to obtuse, membranous to papery;

apex brown to red.

Spikelets

2–4 × 1–2 mm, sometimes absent in deeper water;

proximal scale 1/2 or more of spikelet length;

floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute.

5–10 × 2–4 mm;

proximal scale usually empty, 3–5 mm, usually 1/2 or more as long as spikelet;

floral scales 8–12 per spikelet, lanceolate (to ovate), 3.5–5 × 2–2.5 mm.

Flowers

perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose;

anthers 0.7–1.2 mm.

perianth bristles 6, equal, the longest equaling achene to exceeding tubercle, very slender, spinules dense;

anthers 1.6–3.5 mm.

Achenes

stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X.

medium brown to gray or dark brown, equilaterally trigonous to compressed trigonous, rarely some biconvex, 2–2.7 × 0.7–1.3 mm, apex tapered to a distinct stramineous beak 0.2–0.9 × 0.3–0.6 mm.

Tubers

terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm;

tubers among culm bases straight, narrowly fusiform, 4–5 mm.

Tubercles

0.1–0.2 × 0.15 mm.

0.4–0.5 × 0.3–0.5 mm.

2n

= 10 (Europe).

Eleocharis parvula

Eleocharis suksdorfiana

Phenology Fruiting summer–fall (north) or late winter–fall (far south). Fruiting late spring–summer.
Habitat Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches Bogs, fens, wet meadows, springs, wet gravel near ponds
Elevation 0–600 m (0–2000 ft) (0–)1100–3300 m ((0–)3600–10800 ft)
Distribution
from FNA
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; AB; BC
[BONAP county map]
Discussion

Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Eleocharis suksdorfiana is usually included in E. quinqueflora, it clearly differs qualitatively as given in the key. A collection from hot springs in Ruby Valley, Elko County, Nevada, has stout perianth bristles less than half of the achene length and may represent an undescribed taxon related to E. suksdorfiana. Eleocharis suksdorfiana closely resembles E. rostellata in its achenes, tubercles, culms, and caudices; it differs in the presence of long horizontal rhizomes and the absence of stoloniferous culms. The achenes of E. suksdorfiana are often finely longitudinally ridged, but in E. rostellata they are often rugulose. Specimens from Coconino and Santa Cruz counties, Arizona, are probably E. suksdorfiana but lack achenes so cannot be identified with certainty.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 106. FNA vol. 23, p. 115.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus parvulus, E. pygmaea, S. nanus E. paucifora var. suksdorfiana, E. quinqueflora var. suksdorfiana
Name authority (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) Beauverd: Bull. Soc. Bot. Genève 13: 267. (1922)
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