Eleocharis parvula |
Eleocharis montana |
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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine |
mountain spikerush |
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Habit | Plants perennial, densely tufted or mat-forming; rhizomes mostly hidden by culms and roots, fairly long, 3 mm thick, hard, cortex persistent, longer internodes to 3 mm, scales persistent, ca. 8 mm, membranous, slightly fibrous. | |
Culms | terete, when dry with few to many blunt ridges, 10–70 cm × 0.5–2 mm, soft to firm, internally hollow with complete transverse septa 2–5 mm apart, usually evident externally except in narrowest culms. |
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Leaves | distal leaf sheaths persistent, not splitting, proximally dark red, distally green or brown, slightly callose, papery, apex obtuse, tooth present, 0.3–1 mm. |
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Spikelets | 2–4 × 1–2 mm, sometimes absent in deeper water; proximal scale 1/2 or more of spikelet length; floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute. |
ovoid, 6–21 × 3–4 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale empty or with flower; floral scales appressed in fruit, 100–500+, (15–)30–40 per mm of rachilla, medium brown to colorless, midrib regions greenish to colorless, ovate, 1.5–2.5 × 1–1.5 mm, entire, apex rounded to subacute, carinate in distal part of spikelet. |
Flowers | perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose; anthers 0.7–1.2 mm. |
perianth bristles 6–8, pale brown, medium stout, from less than 1/2 achene length to sometimes slightly exceeding tubercle, retrorsely spinulose; stamens 1; anthers dark yellow to brown, 0.6–1 mm; styles 3-fid or some 2-fid. |
Achenes | stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X. |
falling with scales, dark green or medium brown, obovoid to obpyriform, biconvex or sometimes some compressedtrigonous in same spikelet, lateral angles prominent, abaxial angle absent or evident, not prominent, 0.8–1.1 × 0.7–0.8 mm, neck short or absent, finely cancellate at 10–20X, sometimes finely rugulose, with 15 horizontal ridges in vertical series. |
Tubers | terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm; tubers among culm bases straight, narrowly fusiform, 4–5 mm. |
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Tubercles | 0.1–0.2 × 0.15 mm. |
brown, pyramidal, mostly depressed, 0.2–0.35 × 0.2–0.4 mm. |
2n | = 10 (Europe). |
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Eleocharis parvula |
Eleocharis montana |
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Phenology | Fruiting summer–fall (north) or late winter–fall (far south). | Fruiting winter–fall. |
Habitat | Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches | Fresh temporary or artificial ponds, ditches, burned savannas, swamp margins |
Elevation | 0–600 m (0–2000 ft) | 20–90[–2800] m (100–300[–9200] ft) |
Distribution |
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
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AL; AZ; FL; GA; LA; TX; Mexico; Central America; South America; West Indies |
Discussion | Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxonomy of the septate-culmed species here treated as Eleocharis montana and E. ravenelii should be evaluated. According to H. K. Svenson (1957), the type of E. montana from near Bogotá, Colombia, is the mountain extreme of the species; it has swollen culms with no visible septation. Specimens from Acadia and St. Landry parishes, Louisiana, are intermediate between Eleocharis montana and E. montevidensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 106. | FNA vol. 23, p. 79. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | Scirpus parvulus, E. pygmaea, S. nanus | Scirpus montanus, E. montana var. nodulosa, E. nodulosa |
Name authority | (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) | (Kunth) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 153. (1817) |
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