Eleocharis parvula |
Eleocharis engelmannii |
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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine |
Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
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Culms | 2–40 cm × 0.5–1.5(–2) mm. |
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Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
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Spikelets | 2–4 × 1–2 mm, sometimes absent in deeper water; proximal scale 1/2 or more of spikelet length; floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute. |
lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
Flowers | perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose; anthers 0.7–1.2 mm. |
perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
Achenes | stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X. |
0.9–1.1(–1.5) × 0.7–1.1 mm. |
Tubers | terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm; tubers among culm bases straight, narrowly fusiform, 4–5 mm. |
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Tubercles | 0.1–0.2 × 0.15 mm. |
depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
2n | = 10 (Europe). |
= 10. |
Eleocharis parvula |
Eleocharis engelmannii |
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Phenology | Fruiting summer–fall (north) or late winter–fall (far south). | Fruiting spring–fall. |
Habitat | Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches | Fresh shores, marshes, disturbed places |
Elevation | 0–600 m (0–2000 ft) | 30–2400 m (100–7900 ft) |
Distribution |
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
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AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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Discussion | Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 106. | FNA vol. 23, p. 104. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae |
Sibling taxa | ||
Synonyms | Scirpus parvulus, E. pygmaea, S. nanus | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa |
Name authority | (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) | Steudel: Syn. Pl. Glumac. 2: 79. (1855) |
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