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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine

Britton's spike-rush

Habit Plants usually annual, tufted, often stoloniferous; rhizomes absent.
Culms

ascending, quadrangular, sulcate (not sulcate in Eleocharis sp.

Leaves

distal leaf sheaths persistent or fugaceous, pale brown or green, sometimes streaked red-brown, membranous to translucent, apex acute to acuminate.

Spikelets

2–4 × 1–2 mm, sometimes absent in deeper water;

proximal scale 1/2 or more of spikelet length;

floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute.

basal spikelets absent; often proliferous, ellipsoid to ovoid, terete, 2–10.5 × 1–2.8 mm, apex acute;

proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, slightly longer, often resembling an involucral bract, elliptic or lanceolate, 1.4–2.7 × 0.8–1.2 mm, apex rounded, midrib markedly thickened and broad;

subproximal scale with a flower;

floral scales spiraled, 13–76, 9–13 per mm of rachilla, colorless or pale brown, streaked or mottled pale brown to red-brown, midribs sometimes green, ovate, elliptic, or obovate, 0.8–2.2 × 0.5–1 mm, papery or membranous, midrib evident to prominent, apex rounded.

Flowers

perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose;

anthers 0.7–1.2 mm.

perianth bristles 5–6 or absent, appressed to achene, white, vestigial to shorter than achene, to 0.3 mm;

spinules not evident at 45X;

stamens 2–3;

anthers 0.2–0.45 mm;

styles 3-fid.

Achenes

stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X.

grayish to pale olive or yellowish, often minutely brown-spotted, obovoid, trigonous, angles prominent, 0.45–0.8 × 0.3–0.55 mm, apex constricted proximal to tubercle, smooth or finely pitted at 30X.

Tubers

terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm;

tubers among culm bases straight, narrowly fusiform, 4–5 mm.

Tubercles

0.1–0.2 × 0.15 mm.

gray, green, or yellowish, birettaform, trigonous, 0.05–0.1 × 0.15–0.3 mm.

Aff

.

Brittonii

), 7–37 cm × 0.2–0.6 mm, soft to firm.

2n

= 10 (Europe).

= 10.

Eleocharis parvula

Eleocharis brittonii

Phenology Fruiting summer–fall (north) or late winter–fall (far south). Fruiting mid spring–winter.
Habitat Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches Freshwater, moist terrestrial sites, peaty ponds, swamps
Elevation 0–600 m (0–2000 ft) 0–100 m (0–300 ft)
Distribution
from FNA
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MO; MS; SC; TN; TX
[WildflowerSearch map]
Discussion

Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Preliminary studies suggest that Eleocharis brittonii may be comprised of two entities that may warrant taxonomic recognition, perhaps at the species level. Typical E. brittonii is usually robust, with floral scales pale brown and papery, bristles well developed, and achenes to 0.8 mm, clearly pitted at 30X. It is known from Alabama, Florida, Georgia, Louisiana, Mississippi, New Jersey, and South Carolina. The second variant is usually small with floral scales whitish with red-brown markings, bristles absent or poorly developed, and achenes not more than 0.6 mm, smooth at 30X. We have annotated specimens of this second variant E. sp. aff. brittonii, known from Arkansas, Florida, Georgia, Louisiana, Mississippi, Tennessee, and Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 106. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus parvulus, E. pygmaea, S. nanus E. microcarpa var. brittonii
Name authority (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) Svenson ex Small: Man. S.E. Fl., 164. (1933)
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