Eleocharis parvula |
Eleocharis bicolor |
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dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine |
two-tone spikerush |
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Habit | Plants probably perennial, tufted, not stoloniferous; rhizomes present, ± horizontal, 0.2–0.4 mm thick, firm, longer internodes 3–7 mm, scales translucent or not. | |
Culms | often ascending, often spotted or streaked red-brown, cross section quadrangular or broadly elliptic, often sulcate, 1.5–8(–25) cm × 0.1–0.3 mm, soft. |
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Leaves | distal leaf sheaths persistent or fugaceous, closely sheathing, not conspicuously wrinkled, colorless or pale brown, sometimes spotted red-brown, delicately membranous, apex narrowly acute. |
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Spikelets | 2–4 × 1–2 mm, sometimes absent in deeper water; proximal scale 1/2 or more of spikelet length; floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute. |
basal spikelets absent; never proliferous, ovoid or ellipsoid, laterally compressed or terete, 1.5–3.7 × 0.8–3 mm, apex acute; proximal scale empty, persistent, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales distichous (spirodistichous), 8–22, 6–10 per mm of rachilla, colorless or pale brown, mottled red-brown to purple, ovate, (1–)2 × 1 mm, membranous, midribs green and red-brown, prominent, apex rounded to obtuse. |
Flowers | perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose; anthers 0.7–1.2 mm. |
perianth bristles present or absent, (1–)4, colorless to white, vestigial to shorter than achene, 0–0.55 mm; spinules not evident at 45X; stamens (1–)3; anthers 0.35–0.45 mm; styles 2-fid (or 3-fid). |
Achenes | stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X. |
whitish to pale brown, obovoid, biconvex, rarely trigonous, angles prominent, 0.5–0.8 × 0.4–0.6 mm, apex slightly constricted proximal to tubercle, smooth. |
Tubers | terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm; tubers among culm bases straight, narrowly fusiform, 4–5 mm. |
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Tubercles | 0.1–0.2 × 0.15 mm. |
sessile, green or red-brown, pyramidal, birettaform or umbonate, 0.15–0.25 × 0.3–0.4 mm. |
2n | = 10 (Europe). |
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Eleocharis parvula |
Eleocharis bicolor |
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Phenology | Fruiting summer–fall (north) or late winter–fall (far south). | Fruiting summer–late fall. |
Habitat | Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches | Moist banks, freshwater, damp pine barrens |
Elevation | 0–600 m (0–2000 ft) | 0–50 m (0–200 ft) |
Distribution |
AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
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FL; GA; MS; West Indies; Central America (Nicaragua) |
Discussion | Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. The culms of Eleocharis bicolor sometimes have complete transverse septa which are prominent when plants are aquatic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 106. | FNA vol. 23, p. 96. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus parvulus, E. pygmaea, S. nanus | |
Name authority | (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) | Chapman: Fl. South. U.S., 517. (1860) |
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