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black sand spikerush, broad fruit spikerush

Habit Plants perennial, forming dense clumps, often stoloniferous. Plants perennial or sometimes annual, sometimes stoloniferous.
Rhizomes

present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick;

internodes very short, scales disintegrating to fibers, 4 mm, papery.

present or absent, horizontal and long or ascending and caudexlike, 0.3–0.5 mm thick, scales usually evident, without tubers.

Culms

erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm.

terete to angled, sometimes compressed, to 100 cm × 5 mm.

Leaves

distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute.

distal leaf sheaths persistent, rarely disintegrating, membranous to papery.

Spikelets

basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute;

proximal scale empty, amplexicaulous, similar to floral scales;

subproximal scale with a flower;

floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded).

to 40 × 7 mm;

floral scales 5–500+ per spikelet.

Flowers

perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute.

anthers 0.2–2.5 mm;

styles 2-fid or 3-fid.

Stamens

3;

anthers 1.2–1.5 mm.

Styles

3-fid.

Achenes

stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X.

variously colored, biconvex to trigonous, 0.4–2 mm, smooth to markedly sculptured at 10X.

Tubercles

stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm.

usually clearly different from achene apex in color, texture, and form, rarely merging with apex.

Eleocharis pachycarpa

Eleocharis sect. Eleocharis

Phenology Fruiting late spring–summer.
Habitat Fresh shores, streambeds, groundwater seeps
Elevation 100–1400 m (300–4600 ft)
Distribution
from FNA
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
Worldwide
Discussion

H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 140 (38 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 91. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae Cyperaceae > Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Subordinate taxa
Name authority E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) unknown
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