Eleocharis pachycarpa |
Eleocharis sect. Eleocharis |
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black sand spikerush, broad fruit spikerush |
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Habit | Plants perennial, forming dense clumps, often stoloniferous. | Plants perennial or sometimes annual, sometimes stoloniferous. |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
present or absent, horizontal and long or ascending and caudexlike, 0.3–0.5 mm thick, scales usually evident, without tubers. |
Culms | erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
terete to angled, sometimes compressed, to 100 cm × 5 mm. |
Leaves | distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
distal leaf sheaths persistent, rarely disintegrating, membranous to papery. |
Spikelets | basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
to 40 × 7 mm; floral scales 5–500+ per spikelet. |
Flowers | perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
anthers 0.2–2.5 mm; styles 2-fid or 3-fid. |
Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
variously colored, biconvex to trigonous, 0.4–2 mm, smooth to markedly sculptured at 10X. |
Tubercles | stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
usually clearly different from achene apex in color, texture, and form, rarely merging with apex. |
Eleocharis pachycarpa |
Eleocharis sect. Eleocharis |
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Phenology | Fruiting late spring–summer. | |
Habitat | Fresh shores, streambeds, groundwater seeps | |
Elevation | 100–1400 m (300–4600 ft) | |
Distribution |
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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Worldwide |
Discussion | H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 140 (38 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 91. | FNA vol. 23. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis |
Sibling taxa | ||
Subordinate taxa | ||
Name authority | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) | unknown |
Web links |