Eleocharis pachycarpa |
Eleocharis flavescens |
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black sand spikerush, broad fruit spikerush |
pale spike-rush, wrinkle-sheath spike-rush, yellow spikerush, yellow spikesedge |
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Habit | Plants perennial, forming dense clumps, often stoloniferous. | Plants with creeping rhizomes 0.5–1 mm thick. | ||||
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
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Culms | erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
3–42 cm × 0.3–0.6 mm. |
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Leaves | distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
distal leaf sheaths often disintegrating, thinly membranous-translucent, inflated distally, often wrinkled, apex blunt. |
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Spikelets | basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
ellipsoid, 1.5–9 × 1–3.5 mm, apex acute to acuminate; proximal scale without flower, not amplexicaulous; floral scales to 65, 5–7 per mm of rachilla, loosely appressed to appressed, elliptic, 1–3 × 0.4–1.6 mm, membranous, apex acute. |
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Flowers | perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
perianth bristles (0–)5–8, typically 7, white to stramineous, spinules dense to few; styles 2-fid, rarely 3-fid. |
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Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
stramineous to green or dark brown, biconvex, rarely trigonous, obovoid to obpryiform, 0.4–1.1 × 0.3–0.8 mm, very finely reticulate at 40X. |
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Tubercles | stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
whitish to stramineous or green, 0.2–0.7 × 0.2–0.4 mm, apex acute to acuminate. |
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Eleocharis pachycarpa |
Eleocharis flavescens |
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Phenology | Fruiting late spring–summer. | |||||
Habitat | Fresh shores, streambeds, groundwater seeps | |||||
Elevation | 100–1400 m (300–4600 ft) | |||||
Distribution |
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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AL; AR; AZ; CA; CT; DE; FL; GA; ID; IL; IN; LA; MA; MD; ME; MI; MN; MS; MT; NC; NH; NJ; NY; OH; PA; RI; SC; TX; UT; VA; VT; WI; WY; NB; NS; ON; QC; South America; temperate North America; West Indies
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Discussion | H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). The varieties of Eleocharis flavescens are difficult to delimit, especially in the south, and identifications of some specimens to variety are problematic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 91. | FNA vol. 23, p. 100. | ||||
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Scirpus flavescens | |||||
Name authority | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) | (Poiret) Urban: Symb. Antill. 4: 116. (1903) | ||||
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