Flora of North America species comparison

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Eleocharis pachycarpa

Eleocharis diandra

black sand spikerush, broad fruit spikerush

Wright's spikerush, Wright's spikesedge
Habit Plants perennial, forming dense clumps, often stoloniferous.
Rhizomes

present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick;

internodes very short, scales disintegrating to fibers, 4 mm, papery.

Culms

erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm.

often spreading or reclining, 2–25 cm × 0.3–1 mm.
Leaves

distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute.

apex of distal leaf sheath acute to acuminate, tooth sometimes present, to 0.2 mm.
Spikelets

basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute;

proximal scale empty, amplexicaulous, similar to floral scales;

subproximal scale with a flower;

floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded).

ovoid, 2–7 × 1–4 mm, apex subacute;

proximal scale either with flower or empty, base encircling 1/2 of culm;

floral scales 50–100, 10 per mm of rachilla, orange to purple-brown, ovate, 1–1.5 × 0.8 mm, midrib slightly keeled, apex rounded to acute.
Flowers

perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute.

perianth bristles absent;

stamens 2(–3);

anthers yellow, 0.2–0.3 mm;

styles 2-fid or some 3-fid.
Stamens

3;

anthers 1.2–1.5 mm.

Styles

3-fid.

Achenes

stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X.

0.7–1 × 0.6–0.9 mm.
Tubercles

stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm.

deltoid 0.1–0.2 × 0.25–0.45 mm, 1/3–1/2 as high as wide, 1/8–1/4 as high and 3/4–9/10 as wide as achene.

Eleocharis pachycarpa

Eleocharis diandra
Phenology Fruiting late spring–summer. Fruiting late summer–fall.
Habitat Fresh shores, streambeds, groundwater seeps Fresh, mostly sandy, shores of large lakes and streams, sometimes slightly tidal
Elevation 100–1400 m (300–4600 ft) 0–100 m (0–300 ft)
Distribution
from FNA
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; MA; NH; NY; VT; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis diandra is close to E. ovata and E. aestuum; it probably should be treated as a distinct species (A. Haines 2001). It is apparently adapted to the greatly fluctuating water levels of rivers and large lakes (e.g., Oneida Lake in New York, Lake Champlain in Vermont). I have not seen specimens of E. diandra from Maine, New Jersey, or Pennsylvania, which may be based on specimens of E. aestuum. Specimens from the Lake-of-the-Woods shore in southwest Ontario are like E. diandra; they have floral scales with apices rounded, not acute as in typical E. diandra. The only recent observations of E. diandra are from the Connecticut River in Massachusetts (1985) and Oneida Lake in New York (1968; A. Haines 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 23, p. 91. FNA vol. 23, p. 103.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) C. Wright: Bull. Torrey Bot. Club 10: 101. (1883)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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