Eleocharis pachycarpa |
Eleocharis baldwinii |
|
---|---|---|
black sand spikerush, broad fruit spikerush |
Baldwin's spike-rush |
|
Habit | Plants perennial, forming dense clumps, often stoloniferous. | Plants annual(?), often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
|
Culms | erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
often arching, sometimes spotted or streaked red-brown, quadrangular or broadly elliptic, sulcate or not, 0.5–25(–30) cm × 0.1–0.4 mm, soft. |
Leaves | distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
distal leaf sheaths persistent or disintegrating, stramineous, green, red-brown, or spotted or streaked red-brown, membranous, translucent, apex obtuse or acute. |
Spikelets | basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
basal spikelets usually present, pistillate; often proliferous, ovoid, ellipsoid, or linear, clearly laterally compressed, 2.3–6.9 × 0.5–2 mm, apex acute, proximal scale empty, amplexicaulous, similar to floral scales (midribs sometimes more prominent and sometimes slightly prolonged beyond lamina); subproximal scale with a flower; floral scales clearly distichous, 3–9, 3–4 per mm of rachilla, yellowish, pale brown, or red-brown, spotted, streaked, or mottled red-brown, narrowly ovate to oblong, (1.5–)2–5 × 0.4–0.6(–1.2) mm, membranous, apex rounded, obtuse or rarely acute, midribs keeled. |
Flowers | perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
perianth bristles 3–7, incurved, colorless, white, or red-brown, shorter than or equaling achene; spinules sparse, retrorse, sharply acute; stamens 3; anthers 1.4–2.2 mm; styles 3-fid. |
Stamens | 3; anthers 1.2–1.5 mm. |
|
Styles | 3-fid. |
|
Achenes | stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
whitish or greenish, spotted dark green, obovoid, trigonous, angles prominent, 0.6–0.9 × 0.4–0.6 mm, apex constricted proximal to tubercle, smooth to finely cancellate at 10–20X. |
Tubercles | stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
pale to red-brown, pyramidal, trigonous, 0.2–0.3(–0.4) × 0.2–0.5 mm. |
Eleocharis pachycarpa |
Eleocharis baldwinii |
|
Phenology | Fruiting late spring–summer. | Fruiting early spring–winter. |
Habitat | Fresh shores, streambeds, groundwater seeps | Freshwater to moist terrestrial sites, pine savannas, cypress ponds, lakeshores |
Elevation | 100–1400 m (300–4600 ft) | 0–60 m (0–200 ft) |
Distribution |
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
|
AL; AR; FL; GA; LA; MS; NC; SC; TX
|
Discussion | H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Coarse plants of Eleocharis baldwinii usually bear more achenes per spikelet, dark colored achenes, culms gray-green when dried, and yellowish sheaths. Finer plants often bear few achenes per spikelet, whitish achenes, culms that are mid green when dried, and reddish sheaths. Achene color may relate to maturity, with darker achenes being fully mature. Although these variants appear to intergrade, they may be recognizable taxonomically. The longer floral scales in the spikelet are usually 3.5– 5 mm. We have not seen vouchers to verify the report of Eleocharis baldwinii from Virginia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 91. | FNA vol. 23, p. 98. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Chaetocyperus baldwinii | |
Name authority | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) | (Torrey) Chapman: Fl. South. U.S., 519. (1860) |
Web links |