Eleocharis pachycarpa |
Eleocharis ambigens |
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black sand spikerush, broad fruit spikerush |
ambiguous spikesedge, éléocharide ambiquë |
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Habit | Plants perennial, forming dense clumps, often stoloniferous. | Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 1–3 cm, scales persistent, 5–10 mm, membranous, not fibrous. |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
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Culms | erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
terete, often with to 16 blunt ridges when dry, 25–80 cm × 0.5–1.5 mm, firm, internally spongy. |
Leaves | distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
distal leaf sheaths persistent, not splitting, proximally dark red, distally red to stramineous or green, often callose, thinly papery (to membranous), apex dark red-brown, obtuse to subtruncate, tooth sometimes present on some culms, to 0.3 mm. |
Spikelets | basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
ovoid to lanceoloid, 5–23 × 2–3(–4) mm, apex acute (to obtuse); proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales often spreading in fruit, 10–60, 3–4 per mm of rachilla, medium brown, rarely red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, in distal part lanceolate, 2.5–3.5 × 1.7 mm, entire, apex acute to rarely obtuse, often carinate in distal part of spikelet. |
Flowers | perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
perianth bristles 2–4(–5), brown, slender to stout, usually very unequal, rudimentary to equaling achene; stamens 3; anthers brown, 1.5–2 mm; styles 2-fid, sometimes some 3-fid. |
Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
not persistent, dark yellow or stramineous, obovoid to obpyriform, all biconvex or some (very rarely all) compressedtrigonous, angles obscure, 1–1.5 × 0.9–1.25 mm, apex rounded, neck absent (to very short), finely rugulose at 10–20X, with 20 or more horizontal ridges in vertical series and/or finely cancellate at 10–30X. |
Tubercles | stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
brown or proximally whitish, pyramidal, depressed, rarely some as high as wide, 0.15–0.3(–0.5) × (0.3–)0.4–0.6 mm. |
2n | = 44, 45, 46. |
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Eleocharis pachycarpa |
Eleocharis ambigens |
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Phenology | Fruiting late spring–summer. | Fruiting spring–summer. |
Habitat | Fresh shores, streambeds, groundwater seeps | Coastal (rarely inland) fresh to brackish pond shores, marshes, disturbed places |
Elevation | 100–1400 m (300–4600 ft) | 0–10 m (0–0 ft) |
Distribution |
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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CT; FL; GA; LA; MA; MS; NC; NJ; RI; SC; TX; VA; QC |
Discussion | H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis ambigens is usually treated as a synonym of E. fallax because some specimens from Massachusetts to Georgia have some 3-fid styles and some obscurely trigonous achenes. A specimen of E. ambigens from Louisiana has trigonous achenes. Eleocharis ambigens is very similar to and perhaps better included in E. uniglumis (S.-O. Strandhede 1967). Intermediates with E. macrostachya occur in Louisiana and Texas. The depressed tubercles are diagnostic of E. ambigens. Eleocharis ambigens is sometimes confused with E. montevidensis, which is readily distinguished by its floral scales with broadly rounded apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 91. | FNA vol. 23, p. 78. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Name authority | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) | Fernald: Rhodora 37: 394. (1935) |
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