Eleocharis obtusa |
Eleocharis parvula |
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blunt spike-rush, blunt spikesedge, broad spike spikerush, egg-shape spike-rush, éléocharide obtuse |
dwarf spikerush, dwarf spikesedge, little-head spike-rush, little-head spikesedge, small spike-rush, éléocharide naine |
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Culms | 3–50(–90) cm × 0.2–2 mm. |
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Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
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Spikelets | broadly ovoid (to ellipsoid or lanceoloid), apex rounded (to acute), (2–)5–13 × (2–)3–4 mm; floral scales 15–150+, 8–20 per mm of rachilla, orange-brown (to stramineous), elliptic, 1.5–2.5 × 1–1.5 mm, midribs seldom keeled, apex broadly rounded. |
2–4 × 1–2 mm, sometimes absent in deeper water; proximal scale 1/2 or more of spikelet length; floral scales 6–10 per spikelet, 1.4–2.7 mm, commonly entirely stramineous, apex rounded to subacute. |
Flowers | perianth bristles (5–)6–7, rarely 0, brown, stout, slightly to usually greatly exceeding tubercle; stamens usually 3; anthers brown to yellow, 0.3–0.6 mm; styles usually 3-fid and 2-fid in same spikelet. |
perianth bristles 6, stramineous, fairly stout to slender, usually equaling achene to slightly exceeding tubercle, sometimes unequal and some 1/2 of achene, very rarely rudimentary, minutely retrorsely spinulose; anthers 0.7–1.2 mm. |
Achenes | 0.9–1.2(–1.3) × 0.7–0.9 mm. |
stramineous, sometimes pale brown, obovoid to obpyriform, thickly trigonous, angles distinct, faces concave to plane, rarely convex, 0.9–1.2 × 0.55–0.75 mm, apex tapered, smooth or faintly rough at 30X. |
Tubercles | deltoid 0.35–0.5 × (0.4–)0.5–0.8 mm, 1/3–2/3 as high as wide, 1/3–1/2 as high and 2/3–9/10 as wide as achene. |
0.1–0.2 × 0.15 mm. |
Tubers | terminating rhizomes usually markedly J- or horseshoe-shaped, body (apart from apical bud) oblong, 2–2.5(–5) × 0.5–1 mm; tubers among culm bases straight, narrowly fusiform, 4–5 mm. |
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2n | = 10. |
= 10 (Europe). |
Eleocharis obtusa |
Eleocharis parvula |
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Phenology | Fruiting summer–fall. | Fruiting summer–fall (north) or late winter–fall (far south). |
Habitat | Fresh shores, marshes, disturbed places | Brackish or saline, mostly coastal tidal marshes, shores, mud flats, swamps, ponds, ditches |
Elevation | 10–1600 m (0–5200 ft) | 0–600 m (0–2000 ft) |
Distribution |
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WA; WI; WV; WY; HI; BC; NS; ON; PE; QC
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AL; AR; CA; CT; FL; GA; IL; IN; KS; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OR; SC; VA; WA; BC; NB; NL; NS; QC; Mexico; Central America (Nicaragua); Eurasia
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Discussion | Extremely uncommon plants of Eleocharis obtusa without perianth bristles may be called E. obtusa var. peasei (type from New Hampshire). Robust plants with distinct caudices, floral scales 2.5 mm, and achenes 1.2–1.3 mm (Eleocharis obtusa var. gigantea Fernald) are rare (specimens seen from the Washington-British Columbia border [type], Arkansas, and the Hawaiian Islands). Dwarf plants (E. obtusa var. jejuna Fernald, type from Maine), with unusually small achenes and floral scales, and tubercles often less than 0.5 mm wide, are occasional in the East and are easily confused with E. ovata and E. aestuum. A few specimens are intermediate with E. engelmannii. Eleocharis obtusa is sometimes treated as conspecific with E. ovata, which consistently differs in its mostly 2-fid styles, mostly two stamens, and especially its narrower tubercles (B. M. H. Larson and P. M. Catling 1996). Eleocharis macounii Fernald has been treated as a synonym of E. obutsa (H. K. Svenson 1957) but is more probably a hybrid between E. intermedia and E. obtusa (P. M. Catling and S. G. Hay 1993; see 34. E. intermedia). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants without well-developed bristles are otherwise typical Eleocharis parvula. S.-O. Strandhede and R. M. T. Dahlgren (1968) provided a detailed description from Scandinavia; the mostly curved tubers of North American plants are differently shaped than the ovoid, mostly nearly straight tubers illustrated by them. Eleocharis parvula is very uncommon inland. Plants lacking spikelets and having rather broad culms with evident aerenchyma (E. parvula forma spongiosa Fassett) that are submerged in tidal zones closely resemble small plants of Sagittaria graminea. Eleocharis parvula has also been reported from North Dakota, South America, and Africa; I have not seen specimens. Plants without achenes or tubers cannot be reliably identified to species. Literature reports from Cuba, Mexico, and Venezuela may be based on specimens of E. coloradoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 105. | FNA vol. 23, p. 106. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Parvulae |
Sibling taxa | ||
Synonyms | Scirpus obtusus, E. obtusa var. ellipsoidales, E. obtusa var. gigantea, E. obtusa var. jejuna, E. obtusa var. peasei | Scirpus parvulus, E. pygmaea, S. nanus |
Name authority | (Willdenow) Schultes: Mant. 2: 89. (1824) | (Roemer & Schultes) Link ex Bluff Nees: Comp. Fl. German. ed. 2, 1: 93. (1836) |
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