Eleocharis montevidensis |
Eleocharis obtusetrigona |
|
---|---|---|
Montevideo spike rush, sand spikerush |
spikerush |
|
Habit | Plants perennial, mat-forming; rhizomes evident, long, 0.7–2 mm thick, firm, cortex persistent, longer internodes 1–2 cm, scales often fugaceous, 6–8 mm, membranous, not fibrous. | Plants perennial; rhizomes 3–4 mm thick, soft to hard, longer internodes 3–8 cm, scales more than 1 cm, tubers absent. |
Culms | terete or cross section elliptic or rectangular, with 5–10 blunt ridges when dry, 25–50 cm × 0.5–1.2 mm, firm to hard, spongy. |
obscurely 3–5-angled to terete, 40–110 cm × (2–)3.6–7.5 mm, soft, not septate-nodulose, internally spongy, transverse septa incomplete; plants never forming filiform, flaccid culms. |
Leaves | distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery, apex usually red-brown, often callose, subtruncate to obtuse, tooth usually present on some or all culms, to 0.9 mm. |
distal leaf sheaths persistent, membranous, apex acute to long-acuminate, sometimes prolonged into a slender bladelike portion to 6 cm. |
Spikelets | usually ovoid or ellipsoid to subcylindric, rarely lanceoloid, 4–12 × (1.5–)2–3 mm, apex rounded to acute; proximal scale amplexicaulous or clasping over 3/4 of culm, entire; subproximal scale empty or with flower; floral scales appressed in fruit, 30–100, 6–10 per mm of rachilla, orange-brown, midrib regions often greenish, oblong to ovate, 1.5–2.5 × 1–1.5 mm, apex entire, broadly rounded, sometimes acute in distal part of spikelet, usually horizontally wrinkled and recurved, often carinate in distal part of spikelet. |
not proliferous, 12–43 × 3.5–6 mm; rachilla joints bearing obscure winglike remnants of floral scales; proximal scale without a flower, amplexicaulous, 3.6–7.5 mm; floral scales 30–125, 1–3 per mm of rachilla, greenish to pale brown, often with pale to dark brown submarginal band or a subapical darker spot, ovate to oblong, 4.3–5.8 × 2.5–3.3 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to acute. |
Flowers | perianth bristles 5–6(–7), stramineous to medium brown, stout, often unequal, much shorter than to equaling achene, rarely all rudimentary; stamens 3; anthers dark yellow to stramineous, 0.8–1.5 mm; styles 3-fid or some 2-fid. |
perianth bristles 6–7, stramineous, margins and spinules reddish to pale brown, stout, flattened, subequal, exceeding achene, 2.5–3.2 mm, coarsely retrorsely spinulose; anthers reddish brown, 1.5–1.9 mm; styles 3-fid or sometimes 2-fid. |
Achenes | falling with scales, dark brown, obovoid to obpyriform, compressed trigonous, angles evident to obscure, 0.7–1 × 0.65–0.8 mm, neck absent to sometimes long, finely rugulose at 10–30X, over 20 horizontal ridges in vertical series, and/or minutely cancellate at 20–30X. |
pale brown, biconvex, obpyriform, 1.7–2.3 × 1.2–1.5 mm, clearly sculptured at 10–15X, each face with 10–15 rows of transversely elongated cells, often isodiametric at achene base, apex usually constricted to neck 0.5–0.6 mm wide. |
Tubercles | brown to whitish, pyramidal, as high as wide or sometimes greatly depressed, (0.1–)0.25–0.3 × 0.2–0.3 mm. |
stramineous, lamelliform to high-pyramidal, 0.8–1.1 × 0.7–0.9 mm. |
2n | = 20. |
|
Eleocharis montevidensis |
Eleocharis obtusetrigona |
|
Phenology | Fruiting spring–fall. | Fruiting spring–fall. |
Habitat | Wet soil, fresh ponds, lakes, streams, springs, seeps, marshes, ditches, grasslands | Fresh, permanent water in ditches |
Elevation | 0–2300 m (0–7500 ft) | 0–20 m (0–100 ft) |
Distribution |
AL; AZ; CA; FL; KS; LA; MS; NC; NM; OK; SC; TX; Mexico; South America
|
TX; e Mexico; South America; Central America (Nicaragua) |
Discussion | The name Eleocharis montana was long misapplied to E. montevidensis. The type of E. palmeri, recognized as a species (H. K. Svenson 1957), is indistinguishable from specimens of E. montevidensis. Eleocharis montevidensis is extremely variable; the broadly rounded, usually wrinkled and recurved floral scales are diagnostic. Achenes often fail to form. Eleocharis montevidensis is apparently closely related to E. parishii, in which the floral scales are less densely placed on the rachilla and acute to rounded, and the spikelets are narrowly lanceoloid to cylindric. In the absence of achenes and rhizomes, some specimens of E. montevidensis are easily mistaken for E. tricostata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In North America Eleocharis obtusetrigona is known from only five collections, two of them originally identified as E. fistulosa (Poiret) Link [= E. acutangula (Roxburgh) Schultes], which is common in Mexico and South America. The original description of E. obtusetrigona is poor and does not clearly distinguish E. obtusangula from E. acutangula. H. K. Svenson (1939) included E. obtusetrigona in E. acutangula, which can be distinguished by its acutely trigonous culms as well as its smaller floral scales and achenes. The tubercles are often spongy as in E. quadrangulata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 80. | FNA vol. 23, p. 118. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Limnochloa |
Sibling taxa | ||
Synonyms | E. arenicola, E. montana subsp. montevidensis, E. palmeri | Limnochloa obtusetrigona, E. fistulosa var. obtusetrigona, E. mutata var. obtusetrigona |
Name authority | Kunth: Enum. Pl. 2: 144. (1837) | (Lindley & Nees) Steudel: Syn. Pl. Glumac. 2: 80. (1855) |
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