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Montevideo spike rush, sand spikerush

decumbent spikerush

Habit Plants perennial, mat-forming; rhizomes evident, long, 0.7–2 mm thick, firm, cortex persistent, longer internodes 1–2 cm, scales often fugaceous, 6–8 mm, membranous, not fibrous. Plants perennial, densely tufted; rhizomes often hidden by culms and roots, fairly long, 3–4 mm thick, hard, cortex persistent, longer internodes from very short to 5 mm, scales usually clearly evident, disintegrating to fibers, 20–25 mm, papery.
Culms

terete or cross section elliptic or rectangular, with 5–10 blunt ridges when dry, 25–50 cm × 0.5–1.2 mm, firm to hard, spongy.

terete, often with 10–18 blunt ridges when dry, 10–50 cm × 0.3–2 mm, firm to rigid, spongy.

Leaves

distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery, apex usually red-brown, often callose, subtruncate to obtuse, tooth usually present on some or all culms, to 0.9 mm.

distal leaf sheaths persistent, not splitting, proximally brown or reddish, distally stramineous, brown, reddish or green, often inflated, papery, apex mostly dark red-brown, subtruncate to obtuse, often callose, tooth absent.

Spikelets

usually ovoid or ellipsoid to subcylindric, rarely lanceoloid, 4–12 × (1.5–)2–3 mm, apex rounded to acute;

proximal scale amplexicaulous or clasping over 3/4 of culm, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 30–100, 6–10 per mm of rachilla, orange-brown, midrib regions often greenish, oblong to ovate, 1.5–2.5 × 1–1.5 mm, apex entire, broadly rounded, sometimes acute in distal part of spikelet, usually horizontally wrinkled and recurved, often carinate in distal part of spikelet.

ovoid, 3–8 × 2–2.5 mm, apex acute;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 10–20, 3 per mm of rachilla, orange-brown, midrib regions often greenish, ovate, 3–3.5 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

Flowers

perianth bristles 5–6(–7), stramineous to medium brown, stout, often unequal, much shorter than to equaling achene, rarely all rudimentary;

stamens 3;

anthers dark yellow to stramineous, 0.8–1.5 mm;

styles 3-fid or some 2-fid.

perianth bristles 6, stramineous, stout, nearly equal, mostly equaling or exceeding tubercle, (0.5–)1–2.2 mm, prominently retrorsely spinulose;

stamens 3(?);

anthers dark yellow to stramineous, 1.2–1.5;

styles 3-fid.

Achenes

falling with scales, dark brown, obovoid to obpyriform, compressed trigonous, angles evident to obscure, 0.7–1 × 0.65–0.8 mm, neck absent to sometimes long, finely rugulose at 10–30X, over 20 horizontal ridges in vertical series, and/or minutely cancellate at 20–30X.

falling with scales, dark brown, obpyriform, nearly eqilaterally- to greatly compressed-trigonous, angles slightly prominent, 1–1.3 × 0.75–0.9 mm, neck absent or short, finely rugulose at 20X with more than 20 horizontal ridges in vertical series, or reticulate or cancellate at 20–30X.

Tubercles

brown to whitish, pyramidal, as high as wide or sometimes greatly depressed, (0.1–)0.25–0.3 × 0.2–0.3 mm.

well developed, whitish, pyramidal, as high as wide to much lower than wide, 0.2–0.6 × 0.4–0.7 mm.

2n

= 20.

Eleocharis montevidensis

Eleocharis decumbens

Phenology Fruiting spring–fall. Fruiting summer–fall.
Habitat Wet soil, fresh ponds, lakes, streams, springs, seeps, marshes, ditches, grasslands Wet fresh meadows, seeps, and lakeshores, in interior montane conifer forests and alpine zones
Elevation 0–2300 m (0–7500 ft) 700–3500 m (2300–11500 ft)
Distribution
from FNA
AL; AZ; CA; FL; KS; LA; MS; NC; NM; OK; SC; TX; Mexico; South America
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[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

The name Eleocharis montana was long misapplied to E. montevidensis. The type of E. palmeri, recognized as a species (H. K. Svenson 1957), is indistinguishable from specimens of E. montevidensis. Eleocharis montevidensis is extremely variable; the broadly rounded, usually wrinkled and recurved floral scales are diagnostic. Achenes often fail to form. Eleocharis montevidensis is apparently closely related to E. parishii, in which the floral scales are less densely placed on the rachilla and acute to rounded, and the spikelets are narrowly lanceoloid to cylindric. In the absence of achenes and rhizomes, some specimens of E. montevidensis are easily mistaken for E. tricostata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis decumbens is known only from Shasta and Tulare counties in California.

Although Eleocharis decumbens has long been ignored or treated as a variety of E. montevidensis, it clearly is a very distinct species, from which it differs especially in its thick rhizomes with fibrous scales and its acute floral scales.

Specimens of Eleocharis decumbens without rhizomes or achenes are easily confused with the apparently closely related E. bolanderi, which often may be distinguished by culms no more than 0.5 mm wide and spikelets with scales no more than 3 mm long. The tubercles of E. decumbens are usually well differentiated from the achenes and about as high as wide; the tubercles of E. bolanderi are often poorly developed and much lower than wide. Three collections from Jackson and Klamath counties in Oregon, lack achenes but are probably E. decumbens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 80. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. arenicola, E. montana subsp. montevidensis, E. palmeri E. montevidensis var. decumbens
Name authority Kunth: Enum. Pl. 2: 144. (1837) C. B. Clarke: Bull. Misc. Inform., addit. ser. 8: 23. (1908)
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