Eleocharis montevidensis |
Eleocharis brittonii |
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Montevideo spike rush, sand spikerush |
Britton's spike-rush |
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Habit | Plants perennial, mat-forming; rhizomes evident, long, 0.7–2 mm thick, firm, cortex persistent, longer internodes 1–2 cm, scales often fugaceous, 6–8 mm, membranous, not fibrous. | Plants usually annual, tufted, often stoloniferous; rhizomes absent. |
Culms | terete or cross section elliptic or rectangular, with 5–10 blunt ridges when dry, 25–50 cm × 0.5–1.2 mm, firm to hard, spongy. |
ascending, quadrangular, sulcate (not sulcate in Eleocharis sp. |
Leaves | distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery, apex usually red-brown, often callose, subtruncate to obtuse, tooth usually present on some or all culms, to 0.9 mm. |
distal leaf sheaths persistent or fugaceous, pale brown or green, sometimes streaked red-brown, membranous to translucent, apex acute to acuminate. |
Spikelets | usually ovoid or ellipsoid to subcylindric, rarely lanceoloid, 4–12 × (1.5–)2–3 mm, apex rounded to acute; proximal scale amplexicaulous or clasping over 3/4 of culm, entire; subproximal scale empty or with flower; floral scales appressed in fruit, 30–100, 6–10 per mm of rachilla, orange-brown, midrib regions often greenish, oblong to ovate, 1.5–2.5 × 1–1.5 mm, apex entire, broadly rounded, sometimes acute in distal part of spikelet, usually horizontally wrinkled and recurved, often carinate in distal part of spikelet. |
basal spikelets absent; often proliferous, ellipsoid to ovoid, terete, 2–10.5 × 1–2.8 mm, apex acute; proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, slightly longer, often resembling an involucral bract, elliptic or lanceolate, 1.4–2.7 × 0.8–1.2 mm, apex rounded, midrib markedly thickened and broad; subproximal scale with a flower; floral scales spiraled, 13–76, 9–13 per mm of rachilla, colorless or pale brown, streaked or mottled pale brown to red-brown, midribs sometimes green, ovate, elliptic, or obovate, 0.8–2.2 × 0.5–1 mm, papery or membranous, midrib evident to prominent, apex rounded. |
Flowers | perianth bristles 5–6(–7), stramineous to medium brown, stout, often unequal, much shorter than to equaling achene, rarely all rudimentary; stamens 3; anthers dark yellow to stramineous, 0.8–1.5 mm; styles 3-fid or some 2-fid. |
perianth bristles 5–6 or absent, appressed to achene, white, vestigial to shorter than achene, to 0.3 mm; spinules not evident at 45X; stamens 2–3; anthers 0.2–0.45 mm; styles 3-fid. |
Achenes | falling with scales, dark brown, obovoid to obpyriform, compressed trigonous, angles evident to obscure, 0.7–1 × 0.65–0.8 mm, neck absent to sometimes long, finely rugulose at 10–30X, over 20 horizontal ridges in vertical series, and/or minutely cancellate at 20–30X. |
grayish to pale olive or yellowish, often minutely brown-spotted, obovoid, trigonous, angles prominent, 0.45–0.8 × 0.3–0.55 mm, apex constricted proximal to tubercle, smooth or finely pitted at 30X. |
Tubercles | brown to whitish, pyramidal, as high as wide or sometimes greatly depressed, (0.1–)0.25–0.3 × 0.2–0.3 mm. |
gray, green, or yellowish, birettaform, trigonous, 0.05–0.1 × 0.15–0.3 mm. |
Aff | . |
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Brittonii | ), 7–37 cm × 0.2–0.6 mm, soft to firm. |
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2n | = 20. |
= 10. |
Eleocharis montevidensis |
Eleocharis brittonii |
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Phenology | Fruiting spring–fall. | Fruiting mid spring–winter. |
Habitat | Wet soil, fresh ponds, lakes, streams, springs, seeps, marshes, ditches, grasslands | Freshwater, moist terrestrial sites, peaty ponds, swamps |
Elevation | 0–2300 m (0–7500 ft) | 0–100 m (0–300 ft) |
Distribution |
AL; AZ; CA; FL; KS; LA; MS; NC; NM; OK; SC; TX; Mexico; South America
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AL; AR; FL; GA; LA; MO; MS; SC; TN; TX
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Discussion | The name Eleocharis montana was long misapplied to E. montevidensis. The type of E. palmeri, recognized as a species (H. K. Svenson 1957), is indistinguishable from specimens of E. montevidensis. Eleocharis montevidensis is extremely variable; the broadly rounded, usually wrinkled and recurved floral scales are diagnostic. Achenes often fail to form. Eleocharis montevidensis is apparently closely related to E. parishii, in which the floral scales are less densely placed on the rachilla and acute to rounded, and the spikelets are narrowly lanceoloid to cylindric. In the absence of achenes and rhizomes, some specimens of E. montevidensis are easily mistaken for E. tricostata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Preliminary studies suggest that Eleocharis brittonii may be comprised of two entities that may warrant taxonomic recognition, perhaps at the species level. Typical E. brittonii is usually robust, with floral scales pale brown and papery, bristles well developed, and achenes to 0.8 mm, clearly pitted at 30X. It is known from Alabama, Florida, Georgia, Louisiana, Mississippi, New Jersey, and South Carolina. The second variant is usually small with floral scales whitish with red-brown markings, bristles absent or poorly developed, and achenes not more than 0.6 mm, smooth at 30X. We have annotated specimens of this second variant E. sp. aff. brittonii, known from Arkansas, Florida, Georgia, Louisiana, Mississippi, Tennessee, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 80. | FNA vol. 23. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. arenicola, E. montana subsp. montevidensis, E. palmeri | E. microcarpa var. brittonii |
Name authority | Kunth: Enum. Pl. 2: 144. (1837) | Svenson ex Small: Man. S.E. Fl., 164. (1933) |
Web links |